Gene Co-expression Modules

Bulk vs. Single-Cell RNA-seq

Author

Brian M. Schilder, Bioinformatician II

library(dplyr) 
library(ggplot2) 
root = "./"
source(file.path(root,"MAIN.R")) 
import_parameters(params)

## **** __Utilized Cores__ **** = 2$subsetGenes
## [1] "protein_coding"
## 
## $subsetCells
## [1] 500
## 
## $resolution
## [1] 0.6
## 
## $resultsPath
## [1] "./Results"
## 
## $nCores
## [1] 2
## 
## $perplexity
## [1] 30

load("./Data/monocle3_CDS.RData") 
nCores <- 4

Graph-autocorrelation Analysis

Find genes of interest through graph-autocorrelation analysis.

# pr_graph_test_res = monocle3::graph_test(cds, neighbor_graph="knn", cores=nCores)
# pr_deg_ids = row.names(subset(pr_graph_test_res, q_value < 0.05))

Louvain Modules

• Identify co-expression modules.
  find_gene_modules essentially runs UMAP on the genes (as opposed to the cells) and then groups them into modules using Louvain community analysis.
• Advantages: Much faster than other alogrithms like WGCNA.
  
• Disadvantages: Seems to be limited of ~1000 genes.

set.seed(2019)
gene_module_df = monocle3::find_gene_modules(cds[var.genes[1:1000], ], 
                                             # resolution=1e-2, 
                                             reduction_method = "UMAP",
                                             max_components = 3,
                                             cores=nCores,
                                             umap.fast_sgd = F, # Slower but determinstic results
                                             random_seed = 2019)
data.table::fwrite(gene_module_df, "./Results/sc.modules_by_gene.txt", sep="\t")
createDT(gene_module_df)

Pheatmap

• Plot the eigengene expression of each gene co-expression module (y-axis) within each cell cluster (x-axis).

# Create pheatmap table
cell_group_df = tibble::tibble(cell=row.names(SummarizedExperiment::colData(cds)),
                               cell_group=monocle3::clusters(cds)[colnames(cds)])
agg_mat = monocle3::aggregate_gene_expression(cds, gene_module_df, cell_group_df)
row.names(agg_mat) = stringr::str_c("Module ", row.names(agg_mat))
colnames(agg_mat) = stringr::str_c("Cluster ", colnames(agg_mat))
module_cluster_exp <- data.frame(as.matrix(agg_mat))
data.table::fwrite(data.table::data.table(module_cluster_exp, keep.rownames = "Module"),
                   "./Results/sc.modules_by_cluster.txt", sep="\t" )
# Plot
pheatmap::pheatmap(agg_mat, cluster_rows=T, cluster_cols=F,
                   scale="column", clustering_method="ward.D2",
                   fontsize=6) 

Differential Module Expression

# module_cluster_exp

3D Scatter

# plotly R documentation: https://plot.ly/r/
# rgl::plot3d(x = gene_module_df$dim_1, y=gene_module_df$dim_2, z=gene_module_df$dim_3, ) 
p.3d <- plotly::plot_ly(gene_module_df, 
                     x = ~dim_1, y = ~dim_2, z = ~dim_3, 
                     color = ~module,  
                     text = ~paste("Gene:",id),
                     colors = "Dark2",  
                     # alpha = .7, 
                     size = 5
                     ) %>%
  plotly::layout(title = 'scRNA-seq Gene Co-expression Modules')  %>%
  plotly::add_markers() %>%
  plotly::layout(scene = list(xaxis = list(title = 'UMAP 1'),
                     yaxis = list(title = 'UMAP 2'),
                     zaxis = list(title = 'UMAP 3')))
htmltools::tagList(setNames(list(p.3d), NULL)) 

Module Expression

monocle3::plot_cells(cds,
                     genes=gene_module_df,
                     group_cells_by="cluster",
                     color_cells_by="cluster",
                     show_trajectory_graph=F) #  rasterize = T (for super high-res figures)

Bulk RNAseq Modules vs. scRNAseq Modules

• Co-expression modules were identified through the WGCNA analysis of bulk monocyte RNA-seq data, conducte by Katia Lopes.
  
• Here we compare the modules identified in the scRNA-seq data here (sc.modules), to the modules identified in the bulk monocyte data (bulk.modules).

# Prepare bulk modules
bulk.modules <- data.table::fread("./Data/Katia_Modules/Gene_modules.tbl.txt") 
# bulk.modules$ensembl.id <- gsub("\\.*","", bulk.modules$gene_ids)
# ensembl.dict <- ENSG_to_HGNC(ensembl_ids = unique(bulk.modules$gene_ids), 
#                              reference_genome="grch38", 
#                              input = "ensembl_transcript_id")
if(!file.exists("./Data/Katia_Modules/ens.geneid.gencode.v30")){
  system("scp schilb03@chimera.hpc.mssm.edu:/sc/hydra/projects/pd-omics/GRCH38.p12/ens.geneid.gencode.v30 ./Data/Katia_Modules")
} 
ensembl.dt <- data.table::fread("./Data/Katia_Modules/ens.geneid.gencode.v30")
ensembl.dict <- setNames(ensembl.dt$GeneSymbol, ensembl.dt$gene_id)
bulk.modules$gene.symbol <- ensembl.dict[bulk.modules$gene_ids]  
bulk.modules <- bulk.modules %>% dplyr::rename(module=moduleColors)
createDT(bulk.modules)

## Warning in instance$preRenderHook(instance): It seems your data is too
## big for client-side DataTables. You may consider server-side processing:
## https://rstudio.github.io/DT/server.html

# Prepare sc modules
sc.modules <- data.table::fread("./Results/sc.modules_by_gene.txt") %>% dplyr::rename(gene.symbol=id)
createDT(sc.modules)

Module-Module Overlap

• Calculate enrichment scores for all combinations of bulk RNAseq and scRNAseq gene modules.
  
• This tells you whether some modules are detectable in both datasets.

genomeSize <- length(union(unique(bulk.modules$gene.symbol), monocle3::fData(cds)$gene_short_name))

MOD.dt <- module_vs_module_enrichment(mod.df1 = bulk.modules, 
                                      mod.df2 = sc.modules, 
                                      genomeSize = genomeSize, 
                                      verbose = F) 

## [1] "mod.df1 contains 66 unique modules."
## [1] "mod.df2 contains 17 unique modules."
## [1] "Conducting enrichment tests on 1122 module-module combinations..."
## [1] "1122 enrichment tests conducted in 1.61 seconds."
## [1] "25 enrichment tests were significant (at FDR ≤ 0.05)."
## [1] "22.73% of mod.df1 modules showed enrichment for a mod.df2 module."
## [1] "82.35% of mod.df2 modules showed enrichment for a mod.df1 module."

createDT(MOD.dt)  

Plot Module-Module Overlap

# library(heatmaply)
mod.cast <- reshape2::acast(MOD.dt, 
                            Module1.name~Module2.name, 
                            value.var="Jaccard.index")
hm <- heatmaply::heatmaply(mod.cast, 
                            k_row = 5, k_col = 5, 
                            height = 10, 
                            column_text_angle = 0, 
                            xlab = "scRNA-seq Louvain Modules",
                            ylab = "bulk RNA-seq WGCNA Modules", 
                            cexRow = .7,
                            key.title = "Jaccard Index") # file = "module-module.heatmap.html"

## Registered S3 method overwritten by 'seriation':
##   method         from 
##   reorder.hclust gclus

# shiny::includeHTML("module-module.heatmap.html")
htmltools::tagList(setNames(list(hm), NULL)) 

Module GO Enrichment

• Iteratively test for enrichment of ontological terms in all modules (both bulk and single-cell).
  
• Take a while for long lists of modules (e.g. the 60+ bulk-RNA-seq modules).

mod.enrich.path <- "./Results/gprofiler2.module.enrichment.txt"
if(file.exists(mod.enrich.path)){
  print(paste("Module GO Enrichment file detected. Importing...",mod.enrich.path))
  mod.enrich <- data.table::fread(mod.enrich.path)
} else {
  bulk.mod.enrich <- gprofiler2.module_enrichment(bulk.modules, dataset = "bulk-RNA-seq")
  sc.mod.enrich <- gprofiler2.module_enrichment(sc.modules, dataset = "sc-RNA-seq")
  # Merge
  mod.enrich <- data.table::rbindlist(list(bulk.mod.enrich, sc.mod.enrich))
  # Correct for multiple tests
  mod.enrich <- mod.enrich %>% dplyr::mutate(FDR = p.adjust(p_value, method="fdr"), 
                                             Bonferroni = p.adjust(p_value, method="bonferroni")) %>% data.frame()
  mod.enrich[mod.enrich == "NULL"] <- NA # is.na(mod.enrich) <- mod.enrich == "NULL" 
  data.table::fwrite(mod.enrich, mod.enrich.path, sep="\t") 
}

## [1] "Module GO Enrichment file detected. Importing... ./Results/gprofiler2.module.enrichment.txt"

mod.enrich.sig <- subset(mod.enrich, Bonferroni <= 0.05)
createDT(head(mod.enrich.sig)) 

Module-Module Similarity: Gene-level vs. Term-level

• Compute how similar each of the top conserved module-module pairs are in terms of GO enrichment.

mm.overlap <- module_vs_module_enrichment.terms(MOD.dt = MOD.dt, 
                                                mod.enrich = mod.enrich.sig,  
                                                top.terms = F,
                                                row.limit = F)

## [1] "Comparing overlap of enrichment terms for all module-module combinations."

createDT(mm.overlap)

Histogram

# Prepare dataframe
jaccard.corr <- merge(x = MOD.dt[,c("Module1.name","Module2.name","Jaccard.index")]  %>%
                        dplyr::rename(Jaccard.index.gene=Jaccard.index), 
                      y = mm.overlap[,c("Module1.name","Module2.name","Jaccard.index", 
                                        "Module1.top.term","Module2.top.term")] %>% 
                        dplyr::rename(Jaccard.index.term=Jaccard.index), 
                      by=c("Module1.name","Module2.name"))
# Remove zeros
# jaccard.corr <- subset(jaccard.corr, Jaccard.index.gene > 0 & Jaccard.index.term > 0)
## Histogram
ggplot(jaccard.corr) + 
  geom_histogram(aes(Jaccard.index.gene, fill="Jaccard.index.gene"), alpha=.5) + 
  geom_histogram(aes(Jaccard.index.term, fill="Jaccard.index.term"), alpha=.5)

## `stat_bin()` using `bins = 30`. Pick better value with `binwidth`.
## `stat_bin()` using `bins = 30`. Pick better value with `binwidth`.

Scatterplot

• See how well module-module similarity scores (Jaccard index) correlate between the gene-level comparisons vs. the term-level comparisons.
  
• Varying how many of the top enrichment terms to use to calculate term-level module-module similarity (e.g. 10, 50, 100, all) did not seem to signficantly impact the correlation between methods (generaly hovered between Pearson’s r = .4-.5).
  
• Label each pair of modules in the scatterplot using the top ontological enrichment terms for each module respectively.
• Jaccard.sum is the sum of both the gene-level and term-level Jaccard similarity scores.
  • This can be used as a way to identify module-module pairs that were similar at multiple levels (and thus have strong evidence of conservation).

## Correlation scatterplot
jaccard.corr <- jaccard.corr %>%
# label.dat <- subset(jaccard.corr, (Jaccard.index.gene >= .05| Jaccard.index.term >=.2) ) %>% 
  dplyr::mutate(Label=paste0("x: ",round(Jaccard.index.gene,2),
                             "\ny: ",round(Jaccard.index.term,2),
                             "\n\n",
                             # Mod 1
                             "Mod1 name: ",Module1.name,
                             "\nTop Term: ",Module1.top.term,
                             "\n\n", 
                             # Mod 2
                             "Mod2 name: ",Module2.name,
                             "\nTop Term: ",Module2.top.term),
                Jaccard.sum = Jaccard.index.gene + Jaccard.index.term)

corr.stat <- cor(jaccard.corr$Jaccard.index.gene, jaccard.corr$Jaccard.index.term, method="pearson")
# Scatterplot
sp <- ggplot(jaccard.corr, aes(x=Jaccard.index.gene, y=Jaccard.index.term, text=Label, color=Jaccard.sum)) + 
  geom_point() +
  # ggrepel::geom_text_repel(data = label.dat, aes(label=Label)) +
  geom_smooth(method=lm, se=T) + 
  labs(title = "Correlation between gene-level vs.term-level module-module Jaccard similarities",
       subtitle = paste("Pearson's r =",round(corr.stat,3)))
ggp <- plotly::ggplotly(sp, tooltip ="text" )
htmltools::tagList(list(ggp)) 

Plot Bulk Hubs on Single-Cell Data

# module-module comparisons
MOD.dt <- data.table::fread("./Results/bulk.modules_vs_sc.modules.txt") %>% 
  arrange(FDR)

MOD.dt

##         Module1.name Module2.name Module1.size Module2.size
## 1              green            2          534          104
## 2               pink            8          236           82
## 3        lightyellow            2          130          104
## 4                red            4          517           66
## 5               pink           14          236           88
## 6           skyblue1           14           35           88
## 7                red           16          517          100
## 8        yellowgreen           11           72           25
## 9        darkmagenta            7          148           78
## 10       darkmagenta           10          148           64
## 11       darkmagenta            6          148           58
## 12       darkmagenta           12          148           58
## 13           sienna3           12           72           58
## 14    lavenderblush3           10           41           64
## 15             plum1           13           69           80
## 16             black           17          243           44
## 17          skyblue3           16           70          100
## 18            purple            3          225           53
## 19     mediumpurple3            7           63           78
## 20             white            6           90           58
## 21       darkmagenta            2          148          104
## 22           sienna3            7           72           78
## 23             green            6          534           58
## 24               red            5          517           24
## 25             white            7           90           78
## 26           sienna3            2           72          104
## 27       lightyellow           15          130           32
## 28             plum2            8           50           82
## 29           sienna3            6           72           58
## 30         steelblue            6           77           58
## 31             green           12          534           58
## 32     mediumpurple3           13           63           80
## 33              cyan           15          169           32
## 34            purple            5          225           24
## 35          skyblue3            9           70           24
## 36       lightyellow           14          130           88
## 37        indianred4            6           32           58
## 38            orange            4           92           66
## 39        lightcoral           10           32           64
## 40        lightpink4            3           43           53
## 41            orange            7           92           78
## 42        lightpink4           12           43           58
## 43            coral1           10           40           64
## 44            grey60           17          920           44
## 45       darkmagenta            3          148           53
## 46            purple           10          225           64
## 47       saddlebrown           16           81          100
## 48            purple            4          225           66
## 49     darkturquoise           13          100           80
## 50             green            7          534           78
## 51      midnightblue            1          165           20
## 52           yellow4           14           38           88
## 53         royalblue            7          117           78
## 54             green           10          534           64
## 55     mediumpurple3           12           63           58
## 56             ivory           10           58           64
## 57       darkmagenta            4          148           66
## 58             ivory            4           58           66
## 59           sienna3            3           72           53
## 60     mediumpurple3            4           63           66
## 61              blue            3         1107           53
## 62             plum1           10           69           64
## 63             ivory            7           58           78
## 64        lightcyan1            7           59           78
## 65           sienna3           10           72           64
## 66              pink            1          236           20
## 67       darkmagenta           15          148           32
## 68           skyblue           12           82           58
## 69       darkmagenta           13          148           80
## 70        orangered4            7           64           78
## 71    darkolivegreen            5          218           24
## 72        firebrick4            5           30           24
## 73            brown4           16           55          100
## 74             green           11          534           25
## 75             white           10           90           64
## 76            purple           12          225           58
## 77          skyblue1            5           35           24
## 78             green           13          534           80
## 79            purple           16          225          100
## 80           yellow4            5           38           24
## 81           yellow4            9           38           24
## 82       darkmagenta           17          148           44
## 83      navajowhite2            5           44           24
## 84      midnightblue           14          165           88
## 85    lavenderblush3           11           41           25
## 86          skyblue1           15           35           32
## 87          thistle2            9           49           24
## 88             plum1            2           69          104
## 89             white           13           90           80
## 90            coral1           15           40           32
## 91         turquoise            4         2317           66
## 92         turquoise           17         2317           44
## 93         turquoise           13         2317           80
## 94         turquoise            2         2317          104
## 95         turquoise            3         2317           53
## 96         turquoise            8         2317           82
## 97         turquoise           14         2317           88
## 98         turquoise            6         2317           58
## 99         turquoise            7         2317           78
## 100        turquoise            5         2317           24
## 101        turquoise           16         2317          100
## 102        turquoise           11         2317           25
## 103        turquoise           15         2317           32
## 104        turquoise           10         2317           64
## 105        turquoise            1         2317           20
## 106        turquoise           12         2317           58
## 107        turquoise            9         2317           24
## 108          bisque4            4          145           66
## 109          bisque4           17          145           44
## 110          bisque4           13          145           80
## 111          bisque4            2          145          104
## 112          bisque4            3          145           53
## 113          bisque4            8          145           82
## 114          bisque4           14          145           88
## 115          bisque4            6          145           58
## 116          bisque4            7          145           78
## 117          bisque4            5          145           24
## 118          bisque4           16          145          100
## 119          bisque4           11          145           25
## 120          bisque4           15          145           32
## 121          bisque4           10          145           64
## 122          bisque4            1          145           20
## 123          bisque4           12          145           58
## 124          bisque4            9          145           24
## 125            brown            4          898           66
## 126            brown           17          898           44
## 127            brown           13          898           80
## 128            brown            2          898          104
## 129            brown            3          898           53
## 130            brown            8          898           82
## 131            brown           14          898           88
## 132            brown            6          898           58
## 133            brown            7          898           78
## 134            brown            5          898           24
## 135            brown           16          898          100
## 136            brown           11          898           25
## 137            brown           15          898           32
## 138            brown           10          898           64
## 139            brown            1          898           20
## 140            brown           12          898           58
## 141            brown            9          898           24
## 142         skyblue3            4           70           66
## 143         skyblue3           17           70           44
## 144         skyblue3           13           70           80
## 145         skyblue3            2           70          104
## 146         skyblue3            3           70           53
## 147         skyblue3            8           70           82
## 148         skyblue3           14           70           88
## 149         skyblue3            6           70           58
## 150         skyblue3            7           70           78
## 151         skyblue3            5           70           24
## 152         skyblue3           11           70           25
## 153         skyblue3           15           70           32
## 154         skyblue3           10           70           64
## 155         skyblue3            1           70           20
## 156         skyblue3           12           70           58
## 157              red           17          517           44
## 158              red           13          517           80
## 159              red            2          517          104
## 160              red            3          517           53
## 161              red            8          517           82
## 162              red           14          517           88
## 163              red            6          517           58
## 164              red            7          517           78
## 165              red           11          517           25
## 166              red           15          517           32
## 167              red           10          517           64
## 168              red            1          517           20
## 169              red           12          517           58
## 170              red            9          517           24
## 171             blue            4         1107           66
## 172             blue           17         1107           44
## 173             blue           13         1107           80
## 174             blue            2         1107          104
## 175             blue            8         1107           82
## 176             blue           14         1107           88
## 177             blue            6         1107           58
## 178             blue            7         1107           78
## 179             blue            5         1107           24
## 180             blue           16         1107          100
## 181             blue           11         1107           25
## 182             blue           15         1107           32
## 183             blue           10         1107           64
## 184             blue            1         1107           20
## 185             blue           12         1107           58
## 186             blue            9         1107           24
## 187      darkorange2            4          128           66
## 188      darkorange2           17          128           44
## 189      darkorange2           13          128           80
## 190      darkorange2            2          128          104
## 191      darkorange2            3          128           53
## 192      darkorange2            8          128           82
## 193      darkorange2           14          128           88
## 194      darkorange2            6          128           58
## 195      darkorange2            7          128           78
## 196      darkorange2            5          128           24
## 197      darkorange2           16          128          100
## 198      darkorange2           11          128           25
## 199      darkorange2           15          128           32
## 200      darkorange2           10          128           64
## 201      darkorange2            1          128           20
## 202      darkorange2           12          128           58
## 203      darkorange2            9          128           24
## 204      greenyellow            4          315           66
## 205      greenyellow           17          315           44
## 206      greenyellow           13          315           80
## 207      greenyellow            2          315          104
## 208      greenyellow            3          315           53
## 209      greenyellow            8          315           82
## 210      greenyellow           14          315           88
## 211      greenyellow            6          315           58
## 212      greenyellow            7          315           78
## 213      greenyellow            5          315           24
## 214      greenyellow           16          315          100
## 215      greenyellow           11          315           25
## 216      greenyellow           15          315           32
## 217      greenyellow           10          315           64
## 218      greenyellow            1          315           20
## 219      greenyellow           12          315           58
## 220      greenyellow            9          315           24
## 221   darkolivegreen            4          218           66
## 222   darkolivegreen           17          218           44
## 223   darkolivegreen           13          218           80
## 224   darkolivegreen            2          218          104
## 225   darkolivegreen            3          218           53
## 226   darkolivegreen            8          218           82
## 227   darkolivegreen           14          218           88
## 228   darkolivegreen            6          218           58
## 229   darkolivegreen            7          218           78
## 230   darkolivegreen           16          218          100
## 231   darkolivegreen           11          218           25
## 232   darkolivegreen           15          218           32
## 233   darkolivegreen           10          218           64
## 234   darkolivegreen            1          218           20
## 235   darkolivegreen           12          218           58
## 236   darkolivegreen            9          218           24
## 237           grey60            4          920           66
## 238           grey60           13          920           80
## 239           grey60            2          920          104
## 240           grey60            3          920           53
## 241           grey60            8          920           82
## 242           grey60           14          920           88
## 243           grey60            6          920           58
## 244           grey60            7          920           78
## 245           grey60            5          920           24
## 246           grey60           16          920          100
## 247           grey60           11          920           25
## 248           grey60           15          920           32
## 249           grey60           10          920           64
## 250           grey60            1          920           20
## 251           grey60           12          920           58
## 252           grey60            9          920           24
## 253       orangered3            4           34           66
## 254       orangered3           17           34           44
## 255       orangered3           13           34           80
## 256       orangered3            2           34          104
## 257       orangered3            3           34           53
## 258       orangered3            8           34           82
## 259       orangered3           14           34           88
## 260       orangered3            6           34           58
## 261       orangered3            7           34           78
## 262       orangered3            5           34           24
## 263       orangered3           16           34          100
## 264       orangered3           11           34           25
## 265       orangered3           15           34           32
## 266       orangered3           10           34           64
## 267       orangered3            1           34           20
## 268       orangered3           12           34           58
## 269       orangered3            9           34           24
## 270            plum1            4           69           66
## 271            plum1           17           69           44
## 272            plum1            3           69           53
## 273            plum1            8           69           82
## 274            plum1           14           69           88
## 275            plum1            6           69           58
## 276            plum1            7           69           78
## 277            plum1            5           69           24
## 278            plum1           16           69          100
## 279            plum1           11           69           25
## 280            plum1           15           69           32
## 281            plum1            1           69           20
## 282            plum1           12           69           58
## 283            plum1            9           69           24
## 284             cyan            4          169           66
## 285             cyan           17          169           44
## 286             cyan           13          169           80
## 287             cyan            2          169          104
## 288             cyan            3          169           53
## 289             cyan            8          169           82
## 290             cyan           14          169           88
## 291             cyan            6          169           58
## 292             cyan            7          169           78
## 293             cyan            5          169           24
## 294             cyan           16          169          100
## 295             cyan           11          169           25
## 296             cyan           10          169           64
## 297             cyan            1          169           20
## 298             cyan           12          169           58
## 299             cyan            9          169           24
## 300             pink            4          236           66
## 301             pink           17          236           44
## 302             pink           13          236           80
## 303             pink            2          236          104
## 304             pink            3          236           53
## 305             pink            6          236           58
## 306             pink            7          236           78
## 307             pink            5          236           24
## 308             pink           16          236          100
## 309             pink           11          236           25
## 310             pink           15          236           32
## 311             pink           10          236           64
## 312             pink           12          236           58
## 313             pink            9          236           24
## 314      yellowgreen            4           72           66
## 315      yellowgreen           17           72           44
## 316      yellowgreen           13           72           80
## 317      yellowgreen            2           72          104
## 318      yellowgreen            3           72           53
## 319      yellowgreen            8           72           82
## 320      yellowgreen           14           72           88
## 321      yellowgreen            6           72           58
## 322      yellowgreen            7           72           78
## 323      yellowgreen            5           72           24
## 324      yellowgreen           16           72          100
## 325      yellowgreen           15           72           32
## 326      yellowgreen           10           72           64
## 327      yellowgreen            1           72           20
## 328      yellowgreen           12           72           58
## 329      yellowgreen            9           72           24
## 330            green            4          534           66
## 331            green           17          534           44
## 332            green            3          534           53
## 333            green            8          534           82
## 334            green           14          534           88
## 335            green            5          534           24
## 336            green           16          534          100
## 337            green           15          534           32
## 338            green            1          534           20
## 339            green            9          534           24
## 340           orange           17           92           44
## 341           orange           13           92           80
## 342           orange            2           92          104
## 343           orange            3           92           53
## 344           orange            8           92           82
## 345           orange           14           92           88
## 346           orange            6           92           58
## 347           orange            5           92           24
## 348           orange           16           92          100
## 349           orange           11           92           25
## 350           orange           15           92           32
## 351           orange           10           92           64
## 352           orange            1           92           20
## 353           orange           12           92           58
## 354           orange            9           92           24
## 355            white            4           90           66
## 356            white           17           90           44
## 357            white            2           90          104
## 358            white            3           90           53
## 359            white            8           90           82
## 360            white           14           90           88
## 361            white            5           90           24
## 362            white           16           90          100
## 363            white           11           90           25
## 364            white           15           90           32
## 365            white            1           90           20
## 366            white           12           90           58
## 367            white            9           90           24
## 368              tan            4          177           66
## 369              tan           17          177           44
## 370              tan           13          177           80
## 371              tan            2          177          104
## 372              tan            3          177           53
## 373              tan            8          177           82
## 374              tan           14          177           88
## 375              tan            6          177           58
## 376              tan            7          177           78
## 377              tan            5          177           24
## 378              tan           16          177          100
## 379              tan           11          177           25
## 380              tan           15          177           32
## 381              tan           10          177           64
## 382              tan            1          177           20
## 383              tan           12          177           58
## 384              tan            9          177           24
## 385      saddlebrown            4           81           66
## 386      saddlebrown           17           81           44
## 387      saddlebrown           13           81           80
## 388      saddlebrown            2           81          104
## 389      saddlebrown            3           81           53
## 390      saddlebrown            8           81           82
## 391      saddlebrown           14           81           88
## 392      saddlebrown            6           81           58
## 393      saddlebrown            7           81           78
## 394      saddlebrown            5           81           24
## 395      saddlebrown           11           81           25
## 396      saddlebrown           15           81           32
## 397      saddlebrown           10           81           64
## 398      saddlebrown            1           81           20
## 399      saddlebrown           12           81           58
## 400      saddlebrown            9           81           24
## 401        darkgreen            4          106           66
## 402        darkgreen           17          106           44
## 403        darkgreen           13          106           80
## 404        darkgreen            2          106          104
## 405        darkgreen            3          106           53
## 406        darkgreen            8          106           82
## 407        darkgreen           14          106           88
## 408        darkgreen            6          106           58
## 409        darkgreen            7          106           78
## 410        darkgreen            5          106           24
## 411        darkgreen           16          106          100
## 412        darkgreen           11          106           25
## 413        darkgreen           15          106           32
## 414        darkgreen           10          106           64
## 415        darkgreen            1          106           20
## 416        darkgreen           12          106           58
## 417        darkgreen            9          106           24
## 418      darkmagenta            8          148           82
## 419      darkmagenta           14          148           88
## 420      darkmagenta            5          148           24
## 421      darkmagenta           16          148          100
## 422      darkmagenta           11          148           25
## 423      darkmagenta            1          148           20
## 424      darkmagenta            9          148           24
## 425         thistle1            4           49           66
## 426         thistle1           17           49           44
## 427         thistle1           13           49           80
## 428         thistle1            2           49          104
## 429         thistle1            3           49           53
## 430         thistle1            8           49           82
## 431         thistle1           14           49           88
## 432         thistle1            6           49           58
## 433         thistle1            7           49           78
## 434         thistle1            5           49           24
## 435         thistle1           16           49          100
## 436         thistle1           11           49           25
## 437         thistle1           15           49           32
## 438         thistle1           10           49           64
## 439         thistle1            1           49           20
## 440         thistle1           12           49           58
## 441         thistle1            9           49           24
## 442       firebrick4            4           30           66
## 443       firebrick4           17           30           44
## 444       firebrick4           13           30           80
## 445       firebrick4            2           30          104
## 446       firebrick4            3           30           53
## 447       firebrick4            8           30           82
## 448       firebrick4           14           30           88
## 449       firebrick4            6           30           58
## 450       firebrick4            7           30           78
## 451       firebrick4           16           30          100
## 452       firebrick4           11           30           25
## 453       firebrick4           15           30           32
## 454       firebrick4           10           30           64
## 455       firebrick4            1           30           20
## 456       firebrick4           12           30           58
## 457       firebrick4            9           30           24
## 458           coral2            4           39           66
## 459           coral2           17           39           44
## 460           coral2           13           39           80
## 461           coral2            2           39          104
## 462           coral2            3           39           53
## 463           coral2            8           39           82
## 464           coral2           14           39           88
## 465           coral2            6           39           58
## 466           coral2            7           39           78
## 467           coral2            5           39           24
## 468           coral2           16           39          100
## 469           coral2           11           39           25
## 470           coral2           15           39           32
## 471           coral2           10           39           64
## 472           coral2            1           39           20
## 473           coral2           12           39           58
## 474           coral2            9           39           24
## 475         darkgrey            4           98           66
## 476         darkgrey           17           98           44
## 477         darkgrey           13           98           80
## 478         darkgrey            2           98          104
## 479         darkgrey            3           98           53
## 480         darkgrey            8           98           82
## 481         darkgrey           14           98           88
## 482         darkgrey            6           98           58
## 483         darkgrey            7           98           78
## 484         darkgrey            5           98           24
## 485         darkgrey           16           98          100
## 486         darkgrey           11           98           25
## 487         darkgrey           15           98           32
## 488         darkgrey           10           98           64
## 489         darkgrey            1           98           20
## 490         darkgrey           12           98           58
## 491         darkgrey            9           98           24
## 492          darkred            4          108           66
## 493          darkred           17          108           44
## 494          darkred           13          108           80
## 495          darkred            2          108          104
## 496          darkred            3          108           53
## 497          darkred            8          108           82
## 498          darkred           14          108           88
## 499          darkred            6          108           58
## 500          darkred            7          108           78
## 501          darkred            5          108           24
## 502          darkred           16          108          100
## 503          darkred           11          108           25
## 504          darkred           15          108           32
## 505          darkred           10          108           64
## 506          darkred            1          108           20
## 507          darkred           12          108           58
## 508          darkred            9          108           24
## 509           salmon            4          172           66
## 510           salmon           17          172           44
## 511           salmon           13          172           80
## 512           salmon            2          172          104
## 513           salmon            3          172           53
## 514           salmon            8          172           82
## 515           salmon           14          172           88
## 516           salmon            6          172           58
## 517           salmon            7          172           78
## 518           salmon            5          172           24
## 519           salmon           16          172          100
## 520           salmon           11          172           25
## 521           salmon           15          172           32
## 522           salmon           10          172           64
## 523           salmon            1          172           20
## 524           salmon           12          172           58
## 525           salmon            9          172           24
## 526     navajowhite2            4           44           66
## 527     navajowhite2           17           44           44
## 528     navajowhite2           13           44           80
## 529     navajowhite2            2           44          104
## 530     navajowhite2            3           44           53
## 531     navajowhite2            8           44           82
## 532     navajowhite2           14           44           88
## 533     navajowhite2            6           44           58
## 534     navajowhite2            7           44           78
## 535     navajowhite2           16           44          100
## 536     navajowhite2           11           44           25
## 537     navajowhite2           15           44           32
## 538     navajowhite2           10           44           64
## 539     navajowhite2            1           44           20
## 540     navajowhite2           12           44           58
## 541     navajowhite2            9           44           24
## 542            black            4          243           66
## 543            black           13          243           80
## 544            black            2          243          104
## 545            black            3          243           53
## 546            black            8          243           82
## 547            black           14          243           88
## 548            black            6          243           58
## 549            black            7          243           78
## 550            black            5          243           24
## 551            black           16          243          100
## 552            black           11          243           25
## 553            black           15          243           32
## 554            black           10          243           64
## 555            black            1          243           20
## 556            black           12          243           58
## 557            black            9          243           24
## 558      lightyellow            4          130           66
## 559      lightyellow           17          130           44
## 560      lightyellow           13          130           80
## 561      lightyellow            3          130           53
## 562      lightyellow            8          130           82
## 563      lightyellow            6          130           58
## 564      lightyellow            7          130           78
## 565      lightyellow            5          130           24
## 566      lightyellow           16          130          100
## 567      lightyellow           11          130           25
## 568      lightyellow           10          130           64
## 569      lightyellow            1          130           20
## 570      lightyellow           12          130           58
## 571      lightyellow            9          130           24
## 572         skyblue2            4           39           66
## 573         skyblue2           17           39           44
## 574         skyblue2           13           39           80
## 575         skyblue2            2           39          104
## 576         skyblue2            3           39           53
## 577         skyblue2            8           39           82
## 578         skyblue2           14           39           88
## 579         skyblue2            6           39           58
## 580         skyblue2            7           39           78
## 581         skyblue2            5           39           24
## 582         skyblue2           16           39          100
## 583         skyblue2           11           39           25
## 584         skyblue2           15           39           32
## 585         skyblue2           10           39           64
## 586         skyblue2            1           39           20
## 587         skyblue2           12           39           58
## 588         skyblue2            9           39           24
## 589           purple           17          225           44
## 590           purple           13          225           80
## 591           purple            2          225          104
## 592           purple            8          225           82
## 593           purple           14          225           88
## 594           purple            6          225           58
## 595           purple            7          225           78
## 596           purple           11          225           25
## 597           purple           15          225           32
## 598           purple            1          225           20
## 599           purple            9          225           24
## 600     midnightblue            4          165           66
## 601     midnightblue           17          165           44
## 602     midnightblue           13          165           80
## 603     midnightblue            2          165          104
## 604     midnightblue            3          165           53
## 605     midnightblue            8          165           82
## 606     midnightblue            6          165           58
## 607     midnightblue            7          165           78
## 608     midnightblue            5          165           24
## 609     midnightblue           16          165          100
## 610     midnightblue           11          165           25
## 611     midnightblue           15          165           32
## 612     midnightblue           10          165           64
## 613     midnightblue           12          165           58
## 614     midnightblue            9          165           24
## 615            plum2            4           50           66
## 616            plum2           17           50           44
## 617            plum2           13           50           80
## 618            plum2            2           50          104
## 619            plum2            3           50           53
## 620            plum2           14           50           88
## 621            plum2            6           50           58
## 622            plum2            7           50           78
## 623            plum2            5           50           24
## 624            plum2           16           50          100
## 625            plum2           11           50           25
## 626            plum2           15           50           32
## 627            plum2           10           50           64
## 628            plum2            1           50           20
## 629            plum2           12           50           58
## 630            plum2            9           50           24
## 631    darkturquoise            4          100           66
## 632    darkturquoise           17          100           44
## 633    darkturquoise            2          100          104
## 634    darkturquoise            3          100           53
## 635    darkturquoise            8          100           82
## 636    darkturquoise           14          100           88
## 637    darkturquoise            6          100           58
## 638    darkturquoise            7          100           78
## 639    darkturquoise            5          100           24
## 640    darkturquoise           16          100          100
## 641    darkturquoise           11          100           25
## 642    darkturquoise           15          100           32
## 643    darkturquoise           10          100           64
## 644    darkturquoise            1          100           20
## 645    darkturquoise           12          100           58
## 646    darkturquoise            9          100           24
## 647           coral1            4           40           66
## 648           coral1           17           40           44
## 649           coral1           13           40           80
## 650           coral1            2           40          104
## 651           coral1            3           40           53
## 652           coral1            8           40           82
## 653           coral1           14           40           88
## 654           coral1            6           40           58
## 655           coral1            7           40           78
## 656           coral1            5           40           24
## 657           coral1           16           40          100
## 658           coral1           11           40           25
## 659           coral1            1           40           20
## 660           coral1           12           40           58
## 661           coral1            9           40           24
## 662          skyblue            4           82           66
## 663          skyblue           17           82           44
## 664          skyblue           13           82           80
## 665          skyblue            2           82          104
## 666          skyblue            3           82           53
## 667          skyblue            8           82           82
## 668          skyblue           14           82           88
## 669          skyblue            6           82           58
## 670          skyblue            7           82           78
## 671          skyblue            5           82           24
## 672          skyblue           16           82          100
## 673          skyblue           11           82           25
## 674          skyblue           15           82           32
## 675          skyblue           10           82           64
## 676          skyblue            1           82           20
## 677          skyblue            9           82           24
## 678    darkseagreen4            4           40           66
## 679    darkseagreen4           17           40           44
## 680    darkseagreen4           13           40           80
## 681    darkseagreen4            2           40          104
## 682    darkseagreen4            3           40           53
## 683    darkseagreen4            8           40           82
## 684    darkseagreen4           14           40           88
## 685    darkseagreen4            6           40           58
## 686    darkseagreen4            7           40           78
## 687    darkseagreen4            5           40           24
## 688    darkseagreen4           16           40          100
## 689    darkseagreen4           11           40           25
## 690    darkseagreen4           15           40           32
## 691    darkseagreen4           10           40           64
## 692    darkseagreen4            1           40           20
## 693    darkseagreen4           12           40           58
## 694    darkseagreen4            9           40           24
## 695        steelblue            4           77           66
## 696        steelblue           17           77           44
## 697        steelblue           13           77           80
## 698        steelblue            2           77          104
## 699        steelblue            3           77           53
## 700        steelblue            8           77           82
## 701        steelblue           14           77           88
## 702        steelblue            7           77           78
## 703        steelblue            5           77           24
## 704        steelblue           16           77          100
## 705        steelblue           11           77           25
## 706        steelblue           15           77           32
## 707        steelblue           10           77           64
## 708        steelblue            1           77           20
## 709        steelblue           12           77           58
## 710        steelblue            9           77           24
## 711       lightpink4            4           43           66
## 712       lightpink4           17           43           44
## 713       lightpink4           13           43           80
## 714       lightpink4            2           43          104
## 715       lightpink4            8           43           82
## 716       lightpink4           14           43           88
## 717       lightpink4            6           43           58
## 718       lightpink4            7           43           78
## 719       lightpink4            5           43           24
## 720       lightpink4           16           43          100
## 721       lightpink4           11           43           25
## 722       lightpink4           15           43           32
## 723       lightpink4           10           43           64
## 724       lightpink4            1           43           20
## 725       lightpink4            9           43           24
## 726        royalblue            4          117           66
## 727        royalblue           17          117           44
## 728        royalblue           13          117           80
## 729        royalblue            2          117          104
## 730        royalblue            3          117           53
## 731        royalblue            8          117           82
## 732        royalblue           14          117           88
## 733        royalblue            6          117           58
## 734        royalblue            5          117           24
## 735        royalblue           16          117          100
## 736        royalblue           11          117           25
## 737        royalblue           15          117           32
## 738        royalblue           10          117           64
## 739        royalblue            1          117           20
## 740        royalblue           12          117           58
## 741        royalblue            9          117           24
## 742             plum            4           35           66
## 743             plum           17           35           44
## 744             plum           13           35           80
## 745             plum            2           35          104
## 746             plum            3           35           53
## 747             plum            8           35           82
## 748             plum           14           35           88
## 749             plum            6           35           58
## 750             plum            7           35           78
## 751             plum            5           35           24
## 752             plum           16           35          100
## 753             plum           11           35           25
## 754             plum           15           35           32
## 755             plum           10           35           64
## 756             plum            1           35           20
## 757             plum           12           35           58
## 758             plum            9           35           24
## 759       lightcyan1            4           59           66
## 760       lightcyan1           17           59           44
## 761       lightcyan1           13           59           80
## 762       lightcyan1            2           59          104
## 763       lightcyan1            3           59           53
## 764       lightcyan1            8           59           82
## 765       lightcyan1           14           59           88
## 766       lightcyan1            6           59           58
## 767       lightcyan1            5           59           24
## 768       lightcyan1           16           59          100
## 769       lightcyan1           11           59           25
## 770       lightcyan1           15           59           32
## 771       lightcyan1           10           59           64
## 772       lightcyan1            1           59           20
## 773       lightcyan1           12           59           58
## 774       lightcyan1            9           59           24
## 775            ivory           17           58           44
## 776            ivory           13           58           80
## 777            ivory            2           58          104
## 778            ivory            3           58           53
## 779            ivory            8           58           82
## 780            ivory           14           58           88
## 781            ivory            6           58           58
## 782            ivory            5           58           24
## 783            ivory           16           58          100
## 784            ivory           11           58           25
## 785            ivory           15           58           32
## 786            ivory            1           58           20
## 787            ivory           12           58           58
## 788            ivory            9           58           24
## 789    mediumpurple2            4           33           66
## 790    mediumpurple2           17           33           44
## 791    mediumpurple2           13           33           80
## 792    mediumpurple2            2           33          104
## 793    mediumpurple2            3           33           53
## 794    mediumpurple2            8           33           82
## 795    mediumpurple2           14           33           88
## 796    mediumpurple2            6           33           58
## 797    mediumpurple2            7           33           78
## 798    mediumpurple2            5           33           24
## 799    mediumpurple2           16           33          100
## 800    mediumpurple2           11           33           25
## 801    mediumpurple2           15           33           32
## 802    mediumpurple2           10           33           64
## 803    mediumpurple2            1           33           20
## 804    mediumpurple2           12           33           58
## 805    mediumpurple2            9           33           24
## 806    antiquewhite4            4           39           66
## 807    antiquewhite4           17           39           44
## 808    antiquewhite4           13           39           80
## 809    antiquewhite4            2           39          104
## 810    antiquewhite4            3           39           53
## 811    antiquewhite4            8           39           82
## 812    antiquewhite4           14           39           88
## 813    antiquewhite4            6           39           58
## 814    antiquewhite4            7           39           78
## 815    antiquewhite4            5           39           24
## 816    antiquewhite4           16           39          100
## 817    antiquewhite4           11           39           25
## 818    antiquewhite4           15           39           32
## 819    antiquewhite4           10           39           64
## 820    antiquewhite4            1           39           20
## 821    antiquewhite4           12           39           58
## 822    antiquewhite4            9           39           24
## 823   lavenderblush3            4           41           66
## 824   lavenderblush3           17           41           44
## 825   lavenderblush3           13           41           80
## 826   lavenderblush3            2           41          104
## 827   lavenderblush3            3           41           53
## 828   lavenderblush3            8           41           82
## 829   lavenderblush3           14           41           88
## 830   lavenderblush3            6           41           58
## 831   lavenderblush3            7           41           78
## 832   lavenderblush3            5           41           24
## 833   lavenderblush3           16           41          100
## 834   lavenderblush3           15           41           32
## 835   lavenderblush3            1           41           20
## 836   lavenderblush3           12           41           58
## 837   lavenderblush3            9           41           24
## 838          sienna3            4           72           66
## 839          sienna3           17           72           44
## 840          sienna3           13           72           80
## 841          sienna3            8           72           82
## 842          sienna3           14           72           88
## 843          sienna3            5           72           24
## 844          sienna3           16           72          100
## 845          sienna3           11           72           25
## 846          sienna3           15           72           32
## 847          sienna3            1           72           20
## 848          sienna3            9           72           24
## 849          yellow4            4           38           66
## 850          yellow4           17           38           44
## 851          yellow4           13           38           80
## 852          yellow4            2           38          104
## 853          yellow4            3           38           53
## 854          yellow4            8           38           82
## 855          yellow4            6           38           58
## 856          yellow4            7           38           78
## 857          yellow4           16           38          100
## 858          yellow4           11           38           25
## 859          yellow4           15           38           32
## 860          yellow4           10           38           64
## 861          yellow4            1           38           20
## 862          yellow4           12           38           58
## 863          salmon4            4           46           66
## 864          salmon4           17           46           44
## 865          salmon4           13           46           80
## 866          salmon4            2           46          104
## 867          salmon4            3           46           53
## 868          salmon4            8           46           82
## 869          salmon4           14           46           88
## 870          salmon4            6           46           58
## 871          salmon4            7           46           78
## 872          salmon4            5           46           24
## 873          salmon4           16           46          100
## 874          salmon4           11           46           25
## 875          salmon4           15           46           32
## 876          salmon4           10           46           64
## 877          salmon4            1           46           20
## 878          salmon4           12           46           58
## 879          salmon4            9           46           24
## 880           brown4            4           55           66
## 881           brown4           17           55           44
## 882           brown4           13           55           80
## 883           brown4            2           55          104
## 884           brown4            3           55           53
## 885           brown4            8           55           82
## 886           brown4           14           55           88
## 887           brown4            6           55           58
## 888           brown4            7           55           78
## 889           brown4            5           55           24
## 890           brown4           11           55           25
## 891           brown4           15           55           32
## 892           brown4           10           55           64
## 893           brown4            1           55           20
## 894           brown4           12           55           58
## 895           brown4            9           55           24
## 896   palevioletred3            4           45           66
## 897   palevioletred3           17           45           44
## 898   palevioletred3           13           45           80
## 899   palevioletred3            2           45          104
## 900   palevioletred3            3           45           53
## 901   palevioletred3            8           45           82
## 902   palevioletred3           14           45           88
## 903   palevioletred3            6           45           58
## 904   palevioletred3            7           45           78
## 905   palevioletred3            5           45           24
## 906   palevioletred3           16           45          100
## 907   palevioletred3           11           45           25
## 908   palevioletred3           15           45           32
## 909   palevioletred3           10           45           64
## 910   palevioletred3            1           45           20
## 911   palevioletred3           12           45           58
## 912   palevioletred3            9           45           24
## 913  lightsteelblue1            4           62           66
## 914  lightsteelblue1           17           62           44
## 915  lightsteelblue1           13           62           80
## 916  lightsteelblue1            2           62          104
## 917  lightsteelblue1            3           62           53
## 918  lightsteelblue1            8           62           82
## 919  lightsteelblue1           14           62           88
## 920  lightsteelblue1            6           62           58
## 921  lightsteelblue1            7           62           78
## 922  lightsteelblue1            5           62           24
## 923  lightsteelblue1           16           62          100
## 924  lightsteelblue1           11           62           25
## 925  lightsteelblue1           15           62           32
## 926  lightsteelblue1           10           62           64
## 927  lightsteelblue1            1           62           20
## 928  lightsteelblue1           12           62           58
## 929  lightsteelblue1            9           62           24
## 930    mediumpurple3           17           63           44
## 931    mediumpurple3            2           63          104
## 932    mediumpurple3            3           63           53
## 933    mediumpurple3            8           63           82
## 934    mediumpurple3           14           63           88
## 935    mediumpurple3            6           63           58
## 936    mediumpurple3            5           63           24
## 937    mediumpurple3           16           63          100
## 938    mediumpurple3           11           63           25
## 939    mediumpurple3           15           63           32
## 940    mediumpurple3           10           63           64
## 941    mediumpurple3            1           63           20
## 942    mediumpurple3            9           63           24
## 943      floralwhite            4           56           66
## 944      floralwhite           17           56           44
## 945      floralwhite           13           56           80
## 946      floralwhite            2           56          104
## 947      floralwhite            3           56           53
## 948      floralwhite            8           56           82
## 949      floralwhite           14           56           88
## 950      floralwhite            6           56           58
## 951      floralwhite            7           56           78
## 952      floralwhite            5           56           24
## 953      floralwhite           16           56          100
## 954      floralwhite           11           56           25
## 955      floralwhite           15           56           32
## 956      floralwhite           10           56           64
## 957      floralwhite            1           56           20
## 958      floralwhite           12           56           58
## 959      floralwhite            9           56           24
## 960       orangered4            4           64           66
## 961       orangered4           17           64           44
## 962       orangered4           13           64           80
## 963       orangered4            2           64          104
## 964       orangered4            3           64           53
## 965       orangered4            8           64           82
## 966       orangered4           14           64           88
## 967       orangered4            6           64           58
## 968       orangered4            5           64           24
## 969       orangered4           16           64          100
## 970       orangered4           11           64           25
## 971       orangered4           15           64           32
## 972       orangered4           10           64           64
## 973       orangered4            1           64           20
## 974       orangered4           12           64           58
## 975       orangered4            9           64           24
## 976    darkslateblue            4           51           66
## 977    darkslateblue           17           51           44
## 978    darkslateblue           13           51           80
## 979    darkslateblue            2           51          104
## 980    darkslateblue            3           51           53
## 981    darkslateblue            8           51           82
## 982    darkslateblue           14           51           88
## 983    darkslateblue            6           51           58
## 984    darkslateblue            7           51           78
## 985    darkslateblue            5           51           24
## 986    darkslateblue           16           51          100
## 987    darkslateblue           11           51           25
## 988    darkslateblue           15           51           32
## 989    darkslateblue           10           51           64
## 990    darkslateblue            1           51           20
## 991    darkslateblue           12           51           58
## 992    darkslateblue            9           51           24
## 993        honeydew1            4           41           66
## 994        honeydew1           17           41           44
## 995        honeydew1           13           41           80
## 996        honeydew1            2           41          104
## 997        honeydew1            3           41           53
## 998        honeydew1            8           41           82
## 999        honeydew1           14           41           88
## 1000       honeydew1            6           41           58
## 1001       honeydew1            7           41           78
## 1002       honeydew1            5           41           24
## 1003       honeydew1           16           41          100
## 1004       honeydew1           11           41           25
## 1005       honeydew1           15           41           32
## 1006       honeydew1           10           41           64
## 1007       honeydew1            1           41           20
## 1008       honeydew1           12           41           58
## 1009       honeydew1            9           41           24
## 1010      lightcoral            4           32           66
## 1011      lightcoral           17           32           44
## 1012      lightcoral           13           32           80
## 1013      lightcoral            2           32          104
## 1014      lightcoral            3           32           53
## 1015      lightcoral            8           32           82
## 1016      lightcoral           14           32           88
## 1017      lightcoral            6           32           58
## 1018      lightcoral            7           32           78
## 1019      lightcoral            5           32           24
## 1020      lightcoral           16           32          100
## 1021      lightcoral           11           32           25
## 1022      lightcoral           15           32           32
## 1023      lightcoral            1           32           20
## 1024      lightcoral           12           32           58
## 1025      lightcoral            9           32           24
## 1026  lightsteelblue            4           32           66
## 1027  lightsteelblue           17           32           44
## 1028  lightsteelblue           13           32           80
## 1029  lightsteelblue            2           32          104
## 1030  lightsteelblue            3           32           53
## 1031  lightsteelblue            8           32           82
## 1032  lightsteelblue           14           32           88
## 1033  lightsteelblue            6           32           58
## 1034  lightsteelblue            7           32           78
## 1035  lightsteelblue            5           32           24
## 1036  lightsteelblue           16           32          100
## 1037  lightsteelblue           11           32           25
## 1038  lightsteelblue           15           32           32
## 1039  lightsteelblue           10           32           64
## 1040  lightsteelblue            1           32           20
## 1041  lightsteelblue           12           32           58
## 1042  lightsteelblue            9           32           24
## 1043        thistle2            4           49           66
## 1044        thistle2           17           49           44
## 1045        thistle2           13           49           80
## 1046        thistle2            2           49          104
## 1047        thistle2            3           49           53
## 1048        thistle2            8           49           82
## 1049        thistle2           14           49           88
## 1050        thistle2            6           49           58
## 1051        thistle2            7           49           78
## 1052        thistle2            5           49           24
## 1053        thistle2           16           49          100
## 1054        thistle2           11           49           25
## 1055        thistle2           15           49           32
## 1056        thistle2           10           49           64
## 1057        thistle2            1           49           20
## 1058        thistle2           12           49           58
## 1059      indianred4            4           32           66
## 1060      indianred4           17           32           44
## 1061      indianred4           13           32           80
## 1062      indianred4            2           32          104
## 1063      indianred4            3           32           53
## 1064      indianred4            8           32           82
## 1065      indianred4           14           32           88
## 1066      indianred4            7           32           78
## 1067      indianred4            5           32           24
## 1068      indianred4           16           32          100
## 1069      indianred4           11           32           25
## 1070      indianred4           15           32           32
## 1071      indianred4           10           32           64
## 1072      indianred4            1           32           20
## 1073      indianred4           12           32           58
## 1074      indianred4            9           32           24
## 1075        skyblue1            4           35           66
## 1076        skyblue1           17           35           44
## 1077        skyblue1           13           35           80
## 1078        skyblue1            2           35          104
## 1079        skyblue1            3           35           53
## 1080        skyblue1            8           35           82
## 1081        skyblue1            6           35           58
## 1082        skyblue1            7           35           78
## 1083        skyblue1           16           35          100
## 1084        skyblue1           11           35           25
## 1085        skyblue1           10           35           64
## 1086        skyblue1            1           35           20
## 1087        skyblue1           12           35           58
## 1088        skyblue1            9           35           24
## 1089    mediumorchid            4           39           66
## 1090    mediumorchid           17           39           44
## 1091    mediumorchid           13           39           80
## 1092    mediumorchid            2           39          104
## 1093    mediumorchid            3           39           53
## 1094    mediumorchid            8           39           82
## 1095    mediumorchid           14           39           88
## 1096    mediumorchid            6           39           58
## 1097    mediumorchid            7           39           78
## 1098    mediumorchid            5           39           24
## 1099    mediumorchid           16           39          100
## 1100    mediumorchid           11           39           25
## 1101    mediumorchid           15           39           32
## 1102    mediumorchid           10           39           64
## 1103    mediumorchid            1           39           20
## 1104    mediumorchid           12           39           58
## 1105    mediumorchid            9           39           24
## 1106          maroon            4           44           66
## 1107          maroon           17           44           44
## 1108          maroon           13           44           80
## 1109          maroon            2           44          104
## 1110          maroon            3           44           53
## 1111          maroon            8           44           82
## 1112          maroon           14           44           88
## 1113          maroon            6           44           58
## 1114          maroon            7           44           78
## 1115          maroon            5           44           24
## 1116          maroon           16           44          100
## 1117          maroon           11           44           25
## 1118          maroon           15           44           32
## 1119          maroon           10           44           64
## 1120          maroon            1           44           20
## 1121          maroon           12           44           58
## 1122          maroon            9           44           24
##      Module1.proportion.overlap Module2.proportion.overlap
## 1                         0.103                      0.529
## 2                         0.106                      0.305
## 3                         0.146                      0.183
## 4                         0.044                      0.348
## 5                         0.081                      0.216
## 6                         0.314                      0.125
## 7                         0.048                      0.250
## 8                         0.097                      0.280
## 9                         0.074                      0.141
## 10                        0.054                      0.125
## 11                        0.047                      0.121
## 12                        0.047                      0.121
## 13                        0.069                      0.086
## 14                        0.098                      0.062
## 15                        0.072                      0.062
## 16                        0.025                      0.136
## 17                        0.071                      0.050
## 18                        0.027                      0.113
## 19                        0.063                      0.051
## 20                        0.044                      0.069
## 21                        0.041                      0.058
## 22                        0.056                      0.051
## 23                        0.015                      0.138
## 24                        0.010                      0.208
## 25                        0.044                      0.051
## 26                        0.056                      0.038
## 27                        0.023                      0.094
## 28                        0.060                      0.037
## 29                        0.042                      0.052
## 30                        0.039                      0.052
## 31                        0.013                      0.121
## 32                        0.048                      0.038
## 33                        0.018                      0.094
## 34                        0.013                      0.125
## 35                        0.029                      0.083
## 36                        0.031                      0.045
## 37                        0.062                      0.034
## 38                        0.033                      0.045
## 39                        0.062                      0.031
## 40                        0.047                      0.038
## 41                        0.033                      0.038
## 42                        0.047                      0.034
## 43                        0.050                      0.031
## 44                        0.008                      0.159
## 45                        0.020                      0.057
## 46                        0.018                      0.062
## 47                        0.037                      0.030
## 48                        0.018                      0.061
## 49                        0.030                      0.038
## 50                        0.013                      0.090
## 51                        0.012                      0.100
## 52                        0.053                      0.023
## 53                        0.026                      0.038
## 54                        0.011                      0.094
## 55                        0.032                      0.034
## 56                        0.034                      0.031
## 57                        0.020                      0.045
## 58                        0.034                      0.030
## 59                        0.028                      0.038
## 60                        0.032                      0.030
## 61                        0.007                      0.151
## 62                        0.029                      0.031
## 63                        0.034                      0.026
## 64                        0.034                      0.026
## 65                        0.028                      0.031
## 66                        0.008                      0.100
## 67                        0.014                      0.062
## 68                        0.024                      0.034
## 69                        0.020                      0.038
## 70                        0.031                      0.026
## 71                        0.009                      0.083
## 72                        0.033                      0.042
## 73                        0.036                      0.020
## 74                        0.006                      0.120
## 75                        0.022                      0.031
## 76                        0.013                      0.052
## 77                        0.029                      0.042
## 78                        0.011                      0.075
## 79                        0.018                      0.040
## 80                        0.026                      0.042
## 81                        0.026                      0.042
## 82                        0.014                      0.045
## 83                        0.023                      0.042
## 84                        0.018                      0.034
## 85                        0.024                      0.040
## 86                        0.029                      0.031
## 87                        0.020                      0.042
## 88                        0.029                      0.019
## 89                        0.022                      0.025
## 90                        0.025                      0.031
## 91                        0.002                      0.076
## 92                        0.003                      0.136
## 93                        0.004                      0.112
## 94                        0.002                      0.048
## 95                        0.002                      0.094
## 96                        0.007                      0.195
## 97                        0.003                      0.080
## 98                        0.003                      0.138
## 99                        0.003                      0.090
## 100                       0.000                      0.000
## 101                       0.005                      0.120
## 102                       0.001                      0.080
## 103                       0.001                      0.094
## 104                       0.004                      0.141
## 105                       0.001                      0.100
## 106                       0.003                      0.121
## 107                       0.002                      0.208
## 108                       0.000                      0.000
## 109                       0.000                      0.000
## 110                       0.000                      0.000
## 111                       0.000                      0.000
## 112                       0.000                      0.000
## 113                       0.000                      0.000
## 114                       0.007                      0.011
## 115                       0.000                      0.000
## 116                       0.000                      0.000
## 117                       0.000                      0.000
## 118                       0.007                      0.010
## 119                       0.000                      0.000
## 120                       0.000                      0.000
## 121                       0.007                      0.016
## 122                       0.000                      0.000
## 123                       0.007                      0.017
## 124                       0.000                      0.000
## 125                       0.002                      0.030
## 126                       0.004                      0.091
## 127                       0.004                      0.050
## 128                       0.000                      0.000
## 129                       0.003                      0.057
## 130                       0.004                      0.049
## 131                       0.004                      0.045
## 132                       0.001                      0.017
## 133                       0.001                      0.013
## 134                       0.000                      0.000
## 135                       0.006                      0.050
## 136                       0.003                      0.120
## 137                       0.001                      0.031
## 138                       0.000                      0.000
## 139                       0.000                      0.000
## 140                       0.000                      0.000
## 141                       0.000                      0.000
## 142                       0.014                      0.015
## 143                       0.000                      0.000
## 144                       0.014                      0.012
## 145                       0.000                      0.000
## 146                       0.000                      0.000
## 147                       0.000                      0.000
## 148                       0.000                      0.000
## 149                       0.000                      0.000
## 150                       0.014                      0.013
## 151                       0.000                      0.000
## 152                       0.000                      0.000
## 153                       0.014                      0.031
## 154                       0.000                      0.000
## 155                       0.000                      0.000
## 156                       0.000                      0.000
## 157                       0.000                      0.000
## 158                       0.010                      0.062
## 159                       0.000                      0.000
## 160                       0.008                      0.075
## 161                       0.006                      0.037
## 162                       0.006                      0.034
## 163                       0.002                      0.017
## 164                       0.010                      0.064
## 165                       0.002                      0.040
## 166                       0.004                      0.062
## 167                       0.006                      0.047
## 168                       0.004                      0.100
## 169                       0.004                      0.034
## 170                       0.002                      0.042
## 171                       0.004                      0.061
## 172                       0.000                      0.000
## 173                       0.002                      0.025
## 174                       0.002                      0.019
## 175                       0.005                      0.073
## 176                       0.003                      0.034
## 177                       0.000                      0.000
## 178                       0.004                      0.051
## 179                       0.002                      0.083
## 180                       0.005                      0.060
## 181                       0.002                      0.080
## 182                       0.001                      0.031
## 183                       0.001                      0.016
## 184                       0.000                      0.000
## 185                       0.001                      0.017
## 186                       0.002                      0.083
## 187                       0.000                      0.000
## 188                       0.000                      0.000
## 189                       0.016                      0.025
## 190                       0.000                      0.000
## 191                       0.000                      0.000
## 192                       0.000                      0.000
## 193                       0.016                      0.023
## 194                       0.000                      0.000
## 195                       0.000                      0.000
## 196                       0.000                      0.000
## 197                       0.016                      0.020
## 198                       0.000                      0.000
## 199                       0.000                      0.000
## 200                       0.000                      0.000
## 201                       0.000                      0.000
## 202                       0.000                      0.000
## 203                       0.000                      0.000
## 204                       0.000                      0.000
## 205                       0.006                      0.045
## 206                       0.006                      0.025
## 207                       0.003                      0.010
## 208                       0.000                      0.000
## 209                       0.003                      0.012
## 210                       0.003                      0.011
## 211                       0.000                      0.000
## 212                       0.000                      0.000
## 213                       0.000                      0.000
## 214                       0.003                      0.010
## 215                       0.003                      0.040
## 216                       0.000                      0.000
## 217                       0.003                      0.016
## 218                       0.000                      0.000
## 219                       0.000                      0.000
## 220                       0.000                      0.000
## 221                       0.000                      0.000
## 222                       0.005                      0.023
## 223                       0.000                      0.000
## 224                       0.000                      0.000
## 225                       0.000                      0.000
## 226                       0.000                      0.000
## 227                       0.005                      0.011
## 228                       0.000                      0.000
## 229                       0.000                      0.000
## 230                       0.005                      0.010
## 231                       0.000                      0.000
## 232                       0.005                      0.031
## 233                       0.009                      0.031
## 234                       0.000                      0.000
## 235                       0.000                      0.000
## 236                       0.000                      0.000
## 237                       0.000                      0.000
## 238                       0.007                      0.075
## 239                       0.002                      0.019
## 240                       0.000                      0.000
## 241                       0.007                      0.073
## 242                       0.001                      0.011
## 243                       0.001                      0.017
## 244                       0.002                      0.026
## 245                       0.000                      0.000
## 246                       0.004                      0.040
## 247                       0.000                      0.000
## 248                       0.001                      0.031
## 249                       0.002                      0.031
## 250                       0.000                      0.000
## 251                       0.000                      0.000
## 252                       0.002                      0.083
## 253                       0.000                      0.000
## 254                       0.000                      0.000
## 255                       0.029                      0.012
## 256                       0.000                      0.000
## 257                       0.029                      0.019
## 258                       0.000                      0.000
## 259                       0.000                      0.000
## 260                       0.029                      0.017
## 261                       0.000                      0.000
## 262                       0.000                      0.000
## 263                       0.000                      0.000
## 264                       0.000                      0.000
## 265                       0.000                      0.000
## 266                       0.000                      0.000
## 267                       0.000                      0.000
## 268                       0.029                      0.017
## 269                       0.000                      0.000
## 270                       0.000                      0.000
## 271                       0.014                      0.023
## 272                       0.000                      0.000
## 273                       0.014                      0.012
## 274                       0.014                      0.011
## 275                       0.014                      0.017
## 276                       0.014                      0.013
## 277                       0.000                      0.000
## 278                       0.000                      0.000
## 279                       0.000                      0.000
## 280                       0.000                      0.000
## 281                       0.000                      0.000
## 282                       0.014                      0.017
## 283                       0.014                      0.042
## 284                       0.006                      0.015
## 285                       0.006                      0.023
## 286                       0.000                      0.000
## 287                       0.000                      0.000
## 288                       0.000                      0.000
## 289                       0.000                      0.000
## 290                       0.012                      0.023
## 291                       0.000                      0.000
## 292                       0.000                      0.000
## 293                       0.000                      0.000
## 294                       0.006                      0.010
## 295                       0.000                      0.000
## 296                       0.006                      0.016
## 297                       0.000                      0.000
## 298                       0.006                      0.017
## 299                       0.000                      0.000
## 300                       0.000                      0.000
## 301                       0.000                      0.000
## 302                       0.000                      0.000
## 303                       0.004                      0.010
## 304                       0.000                      0.000
## 305                       0.000                      0.000
## 306                       0.000                      0.000
## 307                       0.000                      0.000
## 308                       0.000                      0.000
## 309                       0.000                      0.000
## 310                       0.000                      0.000
## 311                       0.000                      0.000
## 312                       0.000                      0.000
## 313                       0.004                      0.042
## 314                       0.000                      0.000
## 315                       0.000                      0.000
## 316                       0.000                      0.000
## 317                       0.000                      0.000
## 318                       0.000                      0.000
## 319                       0.000                      0.000
## 320                       0.000                      0.000
## 321                       0.000                      0.000
## 322                       0.000                      0.000
## 323                       0.000                      0.000
## 324                       0.000                      0.000
## 325                       0.000                      0.000
## 326                       0.000                      0.000
## 327                       0.000                      0.000
## 328                       0.000                      0.000
## 329                       0.000                      0.000
## 330                       0.006                      0.045
## 331                       0.004                      0.045
## 332                       0.006                      0.057
## 333                       0.004                      0.024
## 334                       0.004                      0.023
## 335                       0.002                      0.042
## 336                       0.006                      0.030
## 337                       0.004                      0.062
## 338                       0.000                      0.000
## 339                       0.002                      0.042
## 340                       0.000                      0.000
## 341                       0.000                      0.000
## 342                       0.000                      0.000
## 343                       0.000                      0.000
## 344                       0.011                      0.012
## 345                       0.011                      0.011
## 346                       0.000                      0.000
## 347                       0.000                      0.000
## 348                       0.000                      0.000
## 349                       0.000                      0.000
## 350                       0.000                      0.000
## 351                       0.000                      0.000
## 352                       0.000                      0.000
## 353                       0.000                      0.000
## 354                       0.000                      0.000
## 355                       0.011                      0.015
## 356                       0.000                      0.000
## 357                       0.000                      0.000
## 358                       0.011                      0.019
## 359                       0.000                      0.000
## 360                       0.000                      0.000
## 361                       0.011                      0.042
## 362                       0.011                      0.010
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## 1014                      0.000                      0.000
## 1015                      0.031                      0.012
## 1016                      0.000                      0.000
## 1017                      0.000                      0.000
## 1018                      0.000                      0.000
## 1019                      0.000                      0.000
## 1020                      0.000                      0.000
## 1021                      0.000                      0.000
## 1022                      0.000                      0.000
## 1023                      0.000                      0.000
## 1024                      0.000                      0.000
## 1025                      0.000                      0.000
## 1026                      0.000                      0.000
## 1027                      0.000                      0.000
## 1028                      0.000                      0.000
## 1029                      0.000                      0.000
## 1030                      0.000                      0.000
## 1031                      0.000                      0.000
## 1032                      0.031                      0.011
## 1033                      0.000                      0.000
## 1034                      0.000                      0.000
## 1035                      0.000                      0.000
## 1036                      0.031                      0.010
## 1037                      0.000                      0.000
## 1038                      0.000                      0.000
## 1039                      0.000                      0.000
## 1040                      0.000                      0.000
## 1041                      0.000                      0.000
## 1042                      0.000                      0.000
## 1043                      0.000                      0.000
## 1044                      0.000                      0.000
## 1045                      0.000                      0.000
## 1046                      0.000                      0.000
## 1047                      0.000                      0.000
## 1048                      0.020                      0.012
## 1049                      0.020                      0.011
## 1050                      0.000                      0.000
## 1051                      0.000                      0.000
## 1052                      0.000                      0.000
## 1053                      0.000                      0.000
## 1054                      0.000                      0.000
## 1055                      0.020                      0.031
## 1056                      0.000                      0.000
## 1057                      0.000                      0.000
## 1058                      0.000                      0.000
## 1059                      0.000                      0.000
## 1060                      0.000                      0.000
## 1061                      0.000                      0.000
## 1062                      0.000                      0.000
## 1063                      0.000                      0.000
## 1064                      0.000                      0.000
## 1065                      0.000                      0.000
## 1066                      0.000                      0.000
## 1067                      0.000                      0.000
## 1068                      0.000                      0.000
## 1069                      0.000                      0.000
## 1070                      0.000                      0.000
## 1071                      0.000                      0.000
## 1072                      0.000                      0.000
## 1073                      0.031                      0.017
## 1074                      0.000                      0.000
## 1075                      0.000                      0.000
## 1076                      0.000                      0.000
## 1077                      0.000                      0.000
## 1078                      0.000                      0.000
## 1079                      0.000                      0.000
## 1080                      0.000                      0.000
## 1081                      0.029                      0.017
## 1082                      0.000                      0.000
## 1083                      0.000                      0.000
## 1084                      0.000                      0.000
## 1085                      0.029                      0.016
## 1086                      0.000                      0.000
## 1087                      0.000                      0.000
## 1088                      0.000                      0.000
## 1089                      0.000                      0.000
## 1090                      0.000                      0.000
## 1091                      0.000                      0.000
## 1092                      0.000                      0.000
## 1093                      0.026                      0.019
## 1094                      0.000                      0.000
## 1095                      0.000                      0.000
## 1096                      0.000                      0.000
## 1097                      0.000                      0.000
## 1098                      0.000                      0.000
## 1099                      0.000                      0.000
## 1100                      0.000                      0.000
## 1101                      0.000                      0.000
## 1102                      0.000                      0.000
## 1103                      0.000                      0.000
## 1104                      0.000                      0.000
## 1105                      0.000                      0.000
## 1106                      0.000                      0.000
## 1107                      0.000                      0.000
## 1108                      0.000                      0.000
## 1109                      0.000                      0.000
## 1110                      0.000                      0.000
## 1111                      0.000                      0.000
## 1112                      0.023                      0.011
## 1113                      0.000                      0.000
## 1114                      0.000                      0.000
## 1115                      0.000                      0.000
## 1116                      0.000                      0.000
## 1117                      0.000                      0.000
## 1118                      0.000                      0.000
## 1119                      0.000                      0.000
## 1120                      0.000                      0.000
## 1121                      0.000                      0.000
## 1122                      0.000                      0.000
##      intersection.size union.size genome.size      p.value odds.ratio
## 1                   55        583       15712 3.069314e-53 35.3810790
## 2                   25        293       15712 2.565009e-26 32.0023289
## 3                   19        215       15712 1.173101e-20 31.1676207
## 4                   23        560       15712 4.552617e-18 16.3933806
## 5                   19        305       15712 3.905751e-17 19.5253864
## 6                   11        112       15712 3.332392e-17 92.2878071
## 7                   25        592       15712 1.176427e-15 10.2390903
## 8                    7         90       15712 1.416247e-11 93.0018609
## 9                   11        215       15712 1.613356e-10 18.5541744
## 10                   8        204       15712 1.452800e-07 15.8128364
## 11                   7        199       15712 1.148091e-06 15.0914967
## 12                   7        199       15712 1.148091e-06 15.0914967
## 13                   5        125       15712 6.658414e-06 21.9170350
## 14                   4        101       15712 2.263874e-05 28.0942562
## 15                   5        144       15712 2.626283e-05 16.2028906
## 16                   6        281       15712 5.552092e-05 10.2766074
## 17                   5        165       15712 8.235818e-05 12.5802302
## 18                   6        272       15712 1.055837e-04  8.9954118
## 19                   4        137       15712 2.680735e-04 14.2610484
## 20                   4        144       15712 3.372407e-04 13.4049884
## 21                   6        246       15712 4.486237e-04  6.6659739
## 22                   4        146       15712 4.476331e-04 12.3653430
## 23                   8        584       15712 7.252677e-04  4.6007044
## 24                   5        536       15712 9.570314e-04  7.7988135
## 25                   4        164       15712 1.039328e-03  9.7672751
## 26                   4        172       15712 1.319513e-03  9.1372014
## 27                   3        159       15712 2.303584e-03 12.6601906
## 28                   3        129       15712 2.250430e-03 12.5805433
## 29                   3        127       15712 2.376667e-03 12.3118488
## 30                   3        132       15712 2.877320e-03 11.4768650
## 31                   7        585       15712 3.376675e-03  3.9392396
## 32                   3        140       15712 4.052360e-03 10.1051844
## 33                   3        198       15712 4.822370e-03  9.6640904
## 34                   3        246       15712 4.696920e-03  9.9480278
## 35                   2         92       15712 5.068764e-03 20.8574584
## 36                   4        214       15712 6.107632e-03  5.8553659
## 37                   2         88       15712 6.186770e-03 18.5820522
## 38                   3        155       15712 6.812551e-03  8.3200652
## 39                   2         94       15712 7.488318e-03 16.7758400
## 40                   2         94       15712 9.228430e-03 14.9268692
## 41                   3        167       15712 1.077093e-02  6.9853815
## 42                   2         99       15712 1.097501e-02 13.5944464
## 43                   2        102       15712 1.153272e-02 13.2368456
## 44                   7        957       15712 1.324161e-02  3.0571191
## 45                   3        198       15712 1.360674e-02  6.4174229
## 46                   4        285       15712 1.331419e-02  4.6531927
## 47                   3        178       15712 1.491737e-02  6.1573683
## 48                   4        287       15712 1.477387e-02  4.5021459
## 49                   3        177       15712 1.443029e-02  6.2377195
## 50                   7        605       15712 1.665669e-02  2.8259033
## 51                   2        183       15712 1.839797e-02 10.5791393
## 52                   2        124       15712 1.913266e-02 10.0635508
## 53                   3        192       15712 2.043472e-02  5.4448183
## 54                   6        592       15712 2.127921e-02  2.9620111
## 55                   2        119       15712 2.264750e-02  9.1260627
## 56                   2        120       15712 2.332337e-02  8.9777619
## 57                   3        211       15712 2.433797e-02  5.0898918
## 58                   2        122       15712 2.469987e-02  8.6958114
## 59                   2        123       15712 2.455030e-02  8.7293416
## 60                   2        127       15712 2.879529e-02  7.9807609
## 61                   8       1152       15712 3.101988e-02  2.3550876
## 62                   2        131       15712 3.217239e-02  7.4989647
## 63                   2        134       15712 3.362367e-02  7.3182147
## 64                   2        135       15712 3.469350e-02  7.1896913
## 65                   2        134       15712 3.478176e-02  7.1768953
## 66                   2        254       15712 3.573535e-02  7.3386103
## 67                   2        178       15712 3.635185e-02  7.0920756
## 68                   2        138       15712 3.683903e-02  6.9487059
## 69                   3        225       15712 3.976719e-02  4.1600933
## 70                   2        140       15712 4.023705e-02  6.6072051
## 71                   2        240       15712 4.329642e-02  6.5095902
## 72                   1         53       15712 4.486472e-02 23.4504209
## 73                   2        153       15712 4.788635e-02  5.9892219
## 74                   3        556       15712 5.158484e-02  3.8916922
## 75                   2        152       15712 5.203199e-02  5.7020941
## 76                   3        280       15712 5.032152e-02  3.7911164
## 77                   1         58       15712 5.215208e-02 19.9934342
## 78                   6        608       15712 5.458208e-02  2.3191791
## 79                   4        321       15712 5.549572e-02  2.9013888
## 80                   1         61       15712 5.649889e-02 18.3742039
## 81                   1         61       15712 5.649889e-02 18.3742039
## 82                   2        190       15712 6.447823e-02  5.0620275
## 83                   1         67       15712 6.513528e-02 15.8050257
## 84                   3        250       15712 6.540891e-02  3.3681189
## 85                   1         65       15712 6.328109e-02 16.2782891
## 86                   1         66       15712 6.894151e-02 14.8327392
## 87                   1         72       15712 7.227434e-02 14.1580238
## 88                   2        171       15712 7.650036e-02  4.5466614
## 89                   2        168       15712 7.695892e-02  4.5276506
## 90                   1         71       15712 7.840642e-02 12.9244443
## 91                   5       2378       15712 9.746017e-01  0.4727486
## 92                   6       2355       15712 6.470575e-01  0.9125752
## 93                   9       2388       15712 8.526897e-01  0.7317979
## 94                   5       2416       15712 9.996846e-01  0.2904619
## 95                   5       2365       15712 9.079110e-01  0.6013639
## 96                  16       2383       15712 1.441493e-01  1.4042535
## 97                   7       2398       15712 9.815686e-01  0.4981111
## 98                   8       2367       15712 6.376773e-01  0.9247171
## 99                   7       2388       15712 9.534114e-01  0.5686910
## 100                  0       2341       15712 1.000000e+00  0.0000000
## 101                 12       2405       15712 8.197040e-01  0.7872526
## 102                  2       2340       15712 9.015430e-01  0.5023007
## 103                  3       2346       15712 8.706369e-01  0.5975499
## 104                  9       2372       15712 6.153025e-01  0.9457954
## 105                  2       2335       15712 8.167347e-01  0.6420612
## 106                  7       2368       15712 7.712285e-01  0.7928811
## 107                  5       2336       15712 2.741666e-01  1.5224395
## 108                  0        211       15712 1.000000e+00  0.0000000
## 109                  0        189       15712 1.000000e+00  0.0000000
## 110                  0        225       15712 1.000000e+00  0.0000000
## 111                  0        249       15712 1.000000e+00  0.0000000
## 112                  0        198       15712 1.000000e+00  0.0000000
## 113                  0        227       15712 1.000000e+00  0.0000000
## 114                  1        232       15712 5.587601e-01  1.2356113
## 115                  0        203       15712 1.000000e+00  0.0000000
## 116                  0        223       15712 1.000000e+00  0.0000000
## 117                  0        169       15712 1.000000e+00  0.0000000
## 118                  1        244       15712 6.054831e-01  1.0850045
## 119                  0        170       15712 1.000000e+00  0.0000000
## 120                  0        177       15712 1.000000e+00  0.0000000
## 121                  1        208       15712 4.482013e-01  1.7089125
## 122                  0        165       15712 1.000000e+00  0.0000000
## 123                  1        202       15712 4.165095e-01  1.8895341
## 124                  0        169       15712 1.000000e+00  0.0000000
## 125                  2        962       15712 8.976561e-01  0.5144409
## 126                  4        938       15712 2.422389e-01  1.6524956
## 127                  4        974       15712 6.780314e-01  0.8676601
## 128                  0       1002       15712 1.000000e+00  0.0000000
## 129                  3        948       15712 5.904136e-01  0.9897660
## 130                  4        976       15712 6.967435e-01  0.8452979
## 131                  4        982       15712 7.481415e-01  0.7846000
## 132                  1        955       15712 9.672815e-01  0.2886365
## 133                  1        975       15712 9.899691e-01  0.2133776
## 134                  0        922       15712 1.000000e+00  0.0000000
## 135                  5        993       15712 6.832429e-01  0.8675118
## 136                  3        920       15712 1.691265e-01  2.2537013
## 137                  1        929       15712 8.481987e-01  0.5316316
## 138                  0        962       15712 1.000000e+00  0.0000000
## 139                  0        918       15712 1.000000e+00  0.0000000
## 140                  0        956       15712 1.000000e+00  0.0000000
## 141                  0        922       15712 1.000000e+00  0.0000000
## 142                  1        135       15712 2.557010e-01  3.4724754
## 143                  0        114       15712 1.000000e+00  0.0000000
## 144                  1        149       15712 3.010072e-01  2.8546821
## 145                  0        174       15712 1.000000e+00  0.0000000
## 146                  0        123       15712 1.000000e+00  0.0000000
## 147                  0        152       15712 1.000000e+00  0.0000000
## 148                  0        158       15712 1.000000e+00  0.0000000
## 149                  0        128       15712 1.000000e+00  0.0000000
## 150                  1        147       15712 2.947054e-01  2.9291863
## 151                  0         94       15712 1.000000e+00  0.0000000
## 152                  0         95       15712 1.000000e+00  0.0000000
## 153                  1        101       15712 1.332686e-01  7.2960198
## 154                  0        134       15712 1.000000e+00  0.0000000
## 155                  0         90       15712 1.000000e+00  0.0000000
## 156                  0        128       15712 1.000000e+00  0.0000000
## 157                  0        561       15712 1.000000e+00  0.0000000
## 158                  5        592       15712 1.233587e-01  1.9686378
## 159                  0        621       15712 1.000000e+00  0.0000000
## 160                  4        566       15712 9.625968e-02  2.4099202
## 161                  3        596       15712 5.092303e-01  1.1167717
## 162                  3        602       15712 5.570687e-01  1.0375487
## 163                  1        574       15712 8.568942e-01  0.5147012
## 164                  5        590       15712 1.140278e-01  2.0228293
## 165                  1        541       15712 5.670394e-01  1.2250291
## 166                  2        547       15712 2.840158e-01  1.9629967
## 167                  3        578       15712 3.524621e-01  1.4479797
## 168                  2        535       15712 1.392815e-01  3.2739575
## 169                  2        573       15712 5.734242e-01  1.0498814
## 170                  1        540       15712 5.522849e-01  1.2783711
## 171                  4       1169       15712 6.925993e-01  0.8506436
## 172                  0       1151       15712 1.000000e+00  0.0000000
## 173                  2       1185       15712 9.797690e-01  0.3371074
## 174                  2       1209       15712 9.956339e-01  0.2573636
## 175                  6       1183       15712 5.229780e-01  1.0418143
## 176                  3       1192       15712 9.525717e-01  0.4642108
## 177                  0       1165       15712 1.000000e+00  0.0000000
## 178                  4       1181       15712 8.087554e-01  0.7121074
## 179                  2       1129       15712 5.119592e-01  1.1997369
## 180                  6       1201       15712 7.153204e-01  0.8412671
## 181                  2       1130       15712 5.341735e-01  1.1474999
## 182                  1       1138       15712 9.037053e-01  0.4250877
## 183                  1       1170       15712 9.907732e-01  0.2087141
## 184                  0       1127       15712 1.000000e+00  0.0000000
## 185                  1       1164       15712 9.856717e-01  0.2307788
## 186                  2       1129       15712 5.119592e-01  1.1997369
## 187                  0        194       15712 1.000000e+00  0.0000000
## 188                  0        172       15712 1.000000e+00  0.0000000
## 189                  2        206       15712 1.384137e-01  3.1550040
## 190                  0        232       15712 1.000000e+00  0.0000000
## 191                  0        181       15712 1.000000e+00  0.0000000
## 192                  0        210       15712 1.000000e+00  0.0000000
## 193                  2        214       15712 1.610044e-01  2.8600823
## 194                  0        186       15712 1.000000e+00  0.0000000
## 195                  0        206       15712 1.000000e+00  0.0000000
## 196                  0        152       15712 1.000000e+00  0.0000000
## 197                  2        226       15712 1.959866e-01  2.5078623
## 198                  0        153       15712 1.000000e+00  0.0000000
## 199                  0        160       15712 1.000000e+00  0.0000000
## 200                  0        192       15712 1.000000e+00  0.0000000
## 201                  0        148       15712 1.000000e+00  0.0000000
## 202                  0        186       15712 1.000000e+00  0.0000000
## 203                  0        152       15712 1.000000e+00  0.0000000
## 204                  0        381       15712 1.000000e+00  0.0000000
## 205                  2        357       15712 2.204613e-01  2.3358601
## 206                  2        393       15712 4.787765e-01  1.2549132
## 207                  1        418       15712 8.791518e-01  0.4728996
## 208                  0        368       15712 1.000000e+00  0.0000000
## 209                  1        396       15712 8.108110e-01  0.6022007
## 210                  1        402       15712 8.325692e-01  0.5604513
## 211                  0        373       15712 1.000000e+00  0.0000000
## 212                  0        393       15712 1.000000e+00  0.0000000
## 213                  0        339       15712 1.000000e+00  0.0000000
## 214                  1        414       15712 8.688849e-01  0.4921348
## 215                  1        339       15712 3.975151e-01  2.0398082
## 216                  0        347       15712 1.000000e+00  0.0000000
## 217                  1        378       15712 7.271313e-01  0.7751599
## 218                  0        335       15712 1.000000e+00  0.0000000
## 219                  0        373       15712 1.000000e+00  0.0000000
## 220                  0        339       15712 1.000000e+00  0.0000000
## 221                  0        284       15712 1.000000e+00  0.0000000
## 222                  1        261       15712 4.596900e-01  1.6557981
## 223                  0        298       15712 1.000000e+00  0.0000000
## 224                  0        322       15712 1.000000e+00  0.0000000
## 225                  0        271       15712 1.000000e+00  0.0000000
## 226                  0        300       15712 1.000000e+00  0.0000000
## 227                  1        305       15712 7.085736e-01  0.8161014
## 228                  0        276       15712 1.000000e+00  0.0000000
## 229                  0        296       15712 1.000000e+00  0.0000000
## 230                  1        317       15712 7.538075e-01  0.7166259
## 231                  0        243       15712 1.000000e+00  0.0000000
## 232                  1        249       15712 3.608049e-01  2.2984467
## 233                  2        280       15712 2.227350e-01  2.3044563
## 234                  0        238       15712 1.000000e+00  0.0000000
## 235                  0        276       15712 1.000000e+00  0.0000000
## 236                  0        242       15712 1.000000e+00  0.0000000
## 237                  0        986       15712 1.000000e+00  0.0000000
## 238                  6        994       15712 3.269304e-01  1.3056095
## 239                  2       1022       15712 9.861538e-01  0.3137825
## 240                  0        973       15712 1.000000e+00  0.0000000
## 241                  6        996       15712 3.480029e-01  1.2710813
## 242                  1       1007       15712 9.951315e-01  0.1839306
## 243                  1        977       15712 9.699886e-01  0.2813070
## 244                  2        996       15712 9.475200e-01  0.4218714
## 245                  0        944       15712 1.000000e+00  0.0000000
## 246                  4       1016       15712 8.444458e-01  0.6684877
## 247                  0        945       15712 1.000000e+00  0.0000000
## 248                  1        951       15712 8.552563e-01  0.5181316
## 249                  2        982       15712 8.957228e-01  0.5176056
## 250                  0        940       15712 1.000000e+00  0.0000000
## 251                  0        978       15712 1.000000e+00  0.0000000
## 252                  2        942       15712 4.142802e-01  1.4626061
## 253                  0        100       15712 1.000000e+00  0.0000000
## 254                  0         78       15712 1.000000e+00  0.0000000
## 255                  1        113       15712 1.594857e-01  5.9815518
## 256                  0        138       15712 1.000000e+00  0.0000000
## 257                  1         86       15712 1.086377e-01  9.1029184
## 258                  0        116       15712 1.000000e+00  0.0000000
## 259                  0        122       15712 1.000000e+00  0.0000000
## 260                  1         91       15712 1.182740e-01  8.3000392
## 261                  0        112       15712 1.000000e+00  0.0000000
## 262                  0         58       15712 1.000000e+00  0.0000000
## 263                  0        134       15712 1.000000e+00  0.0000000
## 264                  0         59       15712 1.000000e+00  0.0000000
## 265                  0         66       15712 1.000000e+00  0.0000000
## 266                  0         98       15712 1.000000e+00  0.0000000
## 267                  0         54       15712 1.000000e+00  0.0000000
## 268                  1         91       15712 1.182740e-01  8.3000392
## 269                  0         58       15712 1.000000e+00  0.0000000
## 270                  0        135       15712 1.000000e+00  0.0000000
## 271                  1        112       15712 1.762760e-01  5.3344737
## 272                  0        122       15712 1.000000e+00  0.0000000
## 273                  1        150       15712 3.036031e-01  2.8249670
## 274                  1        156       15712 3.218494e-01  2.6291780
## 275                  1        126       15712 2.256536e-01  4.0201705
## 276                  1        146       15712 2.911710e-01  2.9724434
## 277                  0         93       15712 1.000000e+00  0.0000000
## 278                  0        169       15712 1.000000e+00  0.0000000
## 279                  0         94       15712 1.000000e+00  0.0000000
## 280                  0        101       15712 1.000000e+00  0.0000000
## 281                  0         89       15712 1.000000e+00  0.0000000
## 282                  1        126       15712 2.256536e-01  4.0201705
## 283                  1         92       15712 1.003116e-01  9.9807595
## 284                  1        234       15712 5.109240e-01  1.4173314
## 285                  1        212       15712 3.790399e-01  2.1455000
## 286                  0        249       15712 1.000000e+00  0.0000000
## 287                  0        273       15712 1.000000e+00  0.0000000
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## 1018                 0        110       15712 1.000000e+00  0.0000000
## 1019                 0         56       15712 1.000000e+00  0.0000000
## 1020                 0        132       15712 1.000000e+00  0.0000000
## 1021                 0         57       15712 1.000000e+00  0.0000000
## 1022                 0         64       15712 1.000000e+00  0.0000000
## 1023                 0         52       15712 1.000000e+00  0.0000000
## 1024                 0         90       15712 1.000000e+00  0.0000000
## 1025                 0         56       15712 1.000000e+00  0.0000000
## 1026                 0         98       15712 1.000000e+00  0.0000000
## 1027                 0         76       15712 1.000000e+00  0.0000000
## 1028                 0        112       15712 1.000000e+00  0.0000000
## 1029                 0        136       15712 1.000000e+00  0.0000000
## 1030                 0         85       15712 1.000000e+00  0.0000000
## 1031                 0        114       15712 1.000000e+00  0.0000000
## 1032                 1        119       15712 1.646529e-01  5.7798093
## 1033                 0         90       15712 1.000000e+00  0.0000000
## 1034                 0        110       15712 1.000000e+00  0.0000000
## 1035                 0         56       15712 1.000000e+00  0.0000000
## 1036                 1        131       15712 1.849610e-01  5.0756141
## 1037                 0         57       15712 1.000000e+00  0.0000000
## 1038                 0         64       15712 1.000000e+00  0.0000000
## 1039                 0         96       15712 1.000000e+00  0.0000000
## 1040                 0         52       15712 1.000000e+00  0.0000000
## 1041                 0         90       15712 1.000000e+00  0.0000000
## 1042                 0         56       15712 1.000000e+00  0.0000000
## 1043                 0        115       15712 1.000000e+00  0.0000000
## 1044                 0         93       15712 1.000000e+00  0.0000000
## 1045                 0        129       15712 1.000000e+00  0.0000000
## 1046                 0        153       15712 1.000000e+00  0.0000000
## 1047                 0        102       15712 1.000000e+00  0.0000000
## 1048                 1        130       15712 2.264703e-01  4.0072760
## 1049                 1        136       15712 2.409091e-01  3.7293506
## 1050                 0        107       15712 1.000000e+00  0.0000000
## 1051                 0        127       15712 1.000000e+00  0.0000000
## 1052                 0         73       15712 1.000000e+00  0.0000000
## 1053                 0        149       15712 1.000000e+00  0.0000000
## 1054                 0         74       15712 1.000000e+00  0.0000000
## 1055                 1         80       15712 9.520887e-02 10.5007622
## 1056                 0        113       15712 1.000000e+00  0.0000000
## 1057                 0         69       15712 1.000000e+00  0.0000000
## 1058                 0        107       15712 1.000000e+00  0.0000000
## 1059                 0         98       15712 1.000000e+00  0.0000000
## 1060                 0         76       15712 1.000000e+00  0.0000000
## 1061                 0        112       15712 1.000000e+00  0.0000000
## 1062                 0        136       15712 1.000000e+00  0.0000000
## 1063                 0         85       15712 1.000000e+00  0.0000000
## 1064                 0        114       15712 1.000000e+00  0.0000000
## 1065                 0        120       15712 1.000000e+00  0.0000000
## 1066                 0        110       15712 1.000000e+00  0.0000000
## 1067                 0         56       15712 1.000000e+00  0.0000000
## 1068                 0        132       15712 1.000000e+00  0.0000000
## 1069                 0         57       15712 1.000000e+00  0.0000000
## 1070                 0         64       15712 1.000000e+00  0.0000000
## 1071                 0         96       15712 1.000000e+00  0.0000000
## 1072                 0         52       15712 1.000000e+00  0.0000000
## 1073                 1         89       15712 1.117144e-01  8.8375721
## 1074                 0         56       15712 1.000000e+00  0.0000000
## 1075                 0        101       15712 1.000000e+00  0.0000000
## 1076                 0         79       15712 1.000000e+00  0.0000000
## 1077                 0        115       15712 1.000000e+00  0.0000000
## 1078                 0        139       15712 1.000000e+00  0.0000000
## 1079                 0         88       15712 1.000000e+00  0.0000000
## 1080                 0        117       15712 1.000000e+00  0.0000000
## 1081                 1         92       15712 1.215359e-01  8.0580135
## 1082                 0        113       15712 1.000000e+00  0.0000000
## 1083                 0        135       15712 1.000000e+00  0.0000000
## 1084                 0         60       15712 1.000000e+00  0.0000000
## 1085                 1         98       15712 1.332567e-01  7.2871883
## 1086                 0         55       15712 1.000000e+00  0.0000000
## 1087                 0         93       15712 1.000000e+00  0.0000000
## 1088                 0         59       15712 1.000000e+00  0.0000000
## 1089                 0        105       15712 1.000000e+00  0.0000000
## 1090                 0         83       15712 1.000000e+00  0.0000000
## 1091                 0        119       15712 1.000000e+00  0.0000000
## 1092                 0        143       15712 1.000000e+00  0.0000000
## 1093                 1         91       15712 1.236045e-01  7.9020299
## 1094                 0        121       15712 1.000000e+00  0.0000000
## 1095                 0        127       15712 1.000000e+00  0.0000000
## 1096                 0         97       15712 1.000000e+00  0.0000000
## 1097                 0        117       15712 1.000000e+00  0.0000000
## 1098                 0         63       15712 1.000000e+00  0.0000000
## 1099                 0        139       15712 1.000000e+00  0.0000000
## 1100                 0         64       15712 1.000000e+00  0.0000000
## 1101                 0         71       15712 1.000000e+00  0.0000000
## 1102                 0        103       15712 1.000000e+00  0.0000000
## 1103                 0         59       15712 1.000000e+00  0.0000000
## 1104                 0         97       15712 1.000000e+00  0.0000000
## 1105                 0         63       15712 1.000000e+00  0.0000000
## 1106                 0        110       15712 1.000000e+00  0.0000000
## 1107                 0         88       15712 1.000000e+00  0.0000000
## 1108                 0        124       15712 1.000000e+00  0.0000000
## 1109                 0        148       15712 1.000000e+00  0.0000000
## 1110                 0         97       15712 1.000000e+00  0.0000000
## 1111                 0        126       15712 1.000000e+00  0.0000000
## 1112                 1        131       15712 2.192250e-01  4.1641127
## 1113                 0        102       15712 1.000000e+00  0.0000000
## 1114                 0        122       15712 1.000000e+00  0.0000000
## 1115                 0         68       15712 1.000000e+00  0.0000000
## 1116                 0        144       15712 1.000000e+00  0.0000000
## 1117                 0         69       15712 1.000000e+00  0.0000000
## 1118                 0         76       15712 1.000000e+00  0.0000000
## 1119                 0        108       15712 1.000000e+00  0.0000000
## 1120                 0         64       15712 1.000000e+00  0.0000000
## 1121                 0        102       15712 1.000000e+00  0.0000000
## 1122                 0         68       15712 1.000000e+00  0.0000000
##      Jaccard.index          FDR   Bonferroni
## 1     0.0943396226 3.443770e-50 3.443770e-50
## 2     0.0853242321 1.438970e-23 2.877940e-23
## 3     0.0883720930 4.387399e-18 1.316220e-17
## 4     0.0410714286 1.277009e-15 5.108036e-15
## 5     0.0622950820 7.303754e-15 4.382252e-14
## 6     0.0982142857 7.303754e-15 3.738944e-14
## 7     0.0422297297 1.885645e-13 1.319951e-12
## 8     0.0777777778 1.986286e-09 1.589029e-08
## 9     0.0511627907 2.011317e-08 1.810185e-07
## 10    0.0392156863 1.630042e-05 1.630042e-04
## 11    0.0351758794 1.073465e-04 1.288158e-03
## 12    0.0351758794 1.073465e-04 1.288158e-03
## 13    0.0400000000 5.746723e-04 7.470740e-03
## 14    0.0396039604 1.814333e-03 2.540067e-02
## 15    0.0347222222 1.964460e-03 2.946689e-02
## 16    0.0213523132 3.893404e-03 6.229447e-02
## 17    0.0303030303 5.435640e-03 9.240588e-02
## 18    0.0220588235 6.581387e-03 1.184650e-01
## 19    0.0291970803 1.583045e-02 3.007785e-01
## 20    0.0277777778 1.891920e-02 3.783841e-01
## 21    0.0243902439 2.287981e-02 5.033558e-01
## 22    0.0273972603 2.287981e-02 5.022444e-01
## 23    0.0136986301 3.538045e-02 8.137503e-01
## 24    0.0093283582 4.474122e-02 1.000000e+00
## 25    0.0243902439 4.664502e-02 1.000000e+00
## 26    0.0232558140 5.694206e-02 1.000000e+00
## 27    0.0188679245 9.195243e-02 1.000000e+00
## 28    0.0232558140 9.195243e-02 1.000000e+00
## 29    0.0236220472 9.195243e-02 1.000000e+00
## 30    0.0227272727 1.076118e-01 1.000000e+00
## 31    0.0119658120 1.222139e-01 1.000000e+00
## 32    0.0214285714 1.420859e-01 1.000000e+00
## 33    0.0151515152 1.591382e-01 1.000000e+00
## 34    0.0121951220 1.591382e-01 1.000000e+00
## 35    0.0217391304 1.624901e-01 1.000000e+00
## 36    0.0186915888 1.876096e-01 1.000000e+00
## 37    0.0227272727 1.876096e-01 1.000000e+00
## 38    0.0193548387 2.011495e-01 1.000000e+00
## 39    0.0212765957 2.154332e-01 1.000000e+00
## 40    0.0212765957 2.588575e-01 1.000000e+00
## 41    0.0179640719 2.931895e-01 1.000000e+00
## 42    0.0202020202 2.931895e-01 1.000000e+00
## 43    0.0196078431 3.009236e-01 1.000000e+00
## 44    0.0073145246 3.318861e-01 1.000000e+00
## 45    0.0151515152 3.318861e-01 1.000000e+00
## 46    0.0140350877 3.318861e-01 1.000000e+00
## 47    0.0168539326 3.415773e-01 1.000000e+00
## 48    0.0139372822 3.415773e-01 1.000000e+00
## 49    0.0169491525 3.415773e-01 1.000000e+00
## 50    0.0115702479 3.737762e-01 1.000000e+00
## 51    0.0109289617 4.047554e-01 1.000000e+00
## 52    0.0161290323 4.128240e-01 1.000000e+00
## 53    0.0156250000 4.325993e-01 1.000000e+00
## 54    0.0101351351 4.421347e-01 1.000000e+00
## 55    0.0168067227 4.620090e-01 1.000000e+00
## 56    0.0166666667 4.673005e-01 1.000000e+00
## 57    0.0142180095 4.697162e-01 1.000000e+00
## 58    0.0163934426 4.697162e-01 1.000000e+00
## 59    0.0162601626 4.697162e-01 1.000000e+00
## 60    0.0157480315 5.384720e-01 1.000000e+00
## 61    0.0069444444 5.705623e-01 1.000000e+00
## 62    0.0152671756 5.822166e-01 1.000000e+00
## 63    0.0149253731 5.988215e-01 1.000000e+00
## 64    0.0148148148 6.003868e-01 1.000000e+00
## 65    0.0149253731 6.003868e-01 1.000000e+00
## 66    0.0078740157 6.075009e-01 1.000000e+00
## 67    0.0112359551 6.078439e-01 1.000000e+00
## 68    0.0144927536 6.078439e-01 1.000000e+00
## 69    0.0133333333 6.449424e-01 1.000000e+00
## 70    0.0142857143 6.449424e-01 1.000000e+00
## 71    0.0083333333 6.842054e-01 1.000000e+00
## 72    0.0188679245 6.991418e-01 1.000000e+00
## 73    0.0130718954 7.360066e-01 1.000000e+00
## 74    0.0053956835 7.599303e-01 1.000000e+00
## 75    0.0131578947 7.599303e-01 1.000000e+00
## 76    0.0107142857 7.599303e-01 1.000000e+00
## 77    0.0172413793 7.599303e-01 1.000000e+00
## 78    0.0098684211 7.826143e-01 1.000000e+00
## 79    0.0124610592 7.826143e-01 1.000000e+00
## 80    0.0163934426 7.826143e-01 1.000000e+00
## 81    0.0163934426 7.826143e-01 1.000000e+00
## 82    0.0105263158 8.633976e-01 1.000000e+00
## 83    0.0149253731 8.633976e-01 1.000000e+00
## 84    0.0120000000 8.633976e-01 1.000000e+00
## 85    0.0153846154 8.633976e-01 1.000000e+00
## 86    0.0151515152 8.994462e-01 1.000000e+00
## 87    0.0138888889 9.320897e-01 1.000000e+00
## 88    0.0116959064 9.702012e-01 1.000000e+00
## 89    0.0119047619 9.702012e-01 1.000000e+00
## 90    0.0140845070 9.774667e-01 1.000000e+00
## 91    0.0021026072 1.000000e+00 1.000000e+00
## 92    0.0025477707 1.000000e+00 1.000000e+00
## 93    0.0037688442 1.000000e+00 1.000000e+00
## 94    0.0020695364 1.000000e+00 1.000000e+00
## 95    0.0021141649 1.000000e+00 1.000000e+00
## 96    0.0067142258 1.000000e+00 1.000000e+00
## 97    0.0029190992 1.000000e+00 1.000000e+00
## 98    0.0033798057 1.000000e+00 1.000000e+00
## 99    0.0029313233 1.000000e+00 1.000000e+00
## 100   0.0000000000 1.000000e+00 1.000000e+00
## 101   0.0049896050 1.000000e+00 1.000000e+00
## 102   0.0008547009 1.000000e+00 1.000000e+00
## 103   0.0012787724 1.000000e+00 1.000000e+00
## 104   0.0037942664 1.000000e+00 1.000000e+00
## 105   0.0008565310 1.000000e+00 1.000000e+00
## 106   0.0029560811 1.000000e+00 1.000000e+00
## 107   0.0021404110 1.000000e+00 1.000000e+00
## 108   0.0000000000 1.000000e+00 1.000000e+00
## 109   0.0000000000 1.000000e+00 1.000000e+00
## 110   0.0000000000 1.000000e+00 1.000000e+00
## 111   0.0000000000 1.000000e+00 1.000000e+00
## 112   0.0000000000 1.000000e+00 1.000000e+00
## 113   0.0000000000 1.000000e+00 1.000000e+00
## 114   0.0043103448 1.000000e+00 1.000000e+00
## 115   0.0000000000 1.000000e+00 1.000000e+00
## 116   0.0000000000 1.000000e+00 1.000000e+00
## 117   0.0000000000 1.000000e+00 1.000000e+00
## 118   0.0040983607 1.000000e+00 1.000000e+00
## 119   0.0000000000 1.000000e+00 1.000000e+00
## 120   0.0000000000 1.000000e+00 1.000000e+00
## 121   0.0048076923 1.000000e+00 1.000000e+00
## 122   0.0000000000 1.000000e+00 1.000000e+00
## 123   0.0049504950 1.000000e+00 1.000000e+00
## 124   0.0000000000 1.000000e+00 1.000000e+00
## 125   0.0020790021 1.000000e+00 1.000000e+00
## 126   0.0042643923 1.000000e+00 1.000000e+00
## 127   0.0041067762 1.000000e+00 1.000000e+00
## 128   0.0000000000 1.000000e+00 1.000000e+00
## 129   0.0031645570 1.000000e+00 1.000000e+00
## 130   0.0040983607 1.000000e+00 1.000000e+00
## 131   0.0040733198 1.000000e+00 1.000000e+00
## 132   0.0010471204 1.000000e+00 1.000000e+00
## 133   0.0010256410 1.000000e+00 1.000000e+00
## 134   0.0000000000 1.000000e+00 1.000000e+00
## 135   0.0050352467 1.000000e+00 1.000000e+00
## 136   0.0032608696 1.000000e+00 1.000000e+00
## 137   0.0010764263 1.000000e+00 1.000000e+00
## 138   0.0000000000 1.000000e+00 1.000000e+00
## 139   0.0000000000 1.000000e+00 1.000000e+00
## 140   0.0000000000 1.000000e+00 1.000000e+00
## 141   0.0000000000 1.000000e+00 1.000000e+00
## 142   0.0074074074 1.000000e+00 1.000000e+00
## 143   0.0000000000 1.000000e+00 1.000000e+00
## 144   0.0067114094 1.000000e+00 1.000000e+00
## 145   0.0000000000 1.000000e+00 1.000000e+00
## 146   0.0000000000 1.000000e+00 1.000000e+00
## 147   0.0000000000 1.000000e+00 1.000000e+00
## 148   0.0000000000 1.000000e+00 1.000000e+00
## 149   0.0000000000 1.000000e+00 1.000000e+00
## 150   0.0068027211 1.000000e+00 1.000000e+00
## 151   0.0000000000 1.000000e+00 1.000000e+00
## 152   0.0000000000 1.000000e+00 1.000000e+00
## 153   0.0099009901 1.000000e+00 1.000000e+00
## 154   0.0000000000 1.000000e+00 1.000000e+00
## 155   0.0000000000 1.000000e+00 1.000000e+00
## 156   0.0000000000 1.000000e+00 1.000000e+00
## 157   0.0000000000 1.000000e+00 1.000000e+00
## 158   0.0084459459 1.000000e+00 1.000000e+00
## 159   0.0000000000 1.000000e+00 1.000000e+00
## 160   0.0070671378 1.000000e+00 1.000000e+00
## 161   0.0050335570 1.000000e+00 1.000000e+00
## 162   0.0049833887 1.000000e+00 1.000000e+00
## 163   0.0017421603 1.000000e+00 1.000000e+00
## 164   0.0084745763 1.000000e+00 1.000000e+00
## 165   0.0018484288 1.000000e+00 1.000000e+00
## 166   0.0036563071 1.000000e+00 1.000000e+00
## 167   0.0051903114 1.000000e+00 1.000000e+00
## 168   0.0037383178 1.000000e+00 1.000000e+00
## 169   0.0034904014 1.000000e+00 1.000000e+00
## 170   0.0018518519 1.000000e+00 1.000000e+00
## 171   0.0034217280 1.000000e+00 1.000000e+00
## 172   0.0000000000 1.000000e+00 1.000000e+00
## 173   0.0016877637 1.000000e+00 1.000000e+00
## 174   0.0016542597 1.000000e+00 1.000000e+00
## 175   0.0050718512 1.000000e+00 1.000000e+00
## 176   0.0025167785 1.000000e+00 1.000000e+00
## 177   0.0000000000 1.000000e+00 1.000000e+00
## 178   0.0033869602 1.000000e+00 1.000000e+00
## 179   0.0017714792 1.000000e+00 1.000000e+00
## 180   0.0049958368 1.000000e+00 1.000000e+00
## 181   0.0017699115 1.000000e+00 1.000000e+00
## 182   0.0008787346 1.000000e+00 1.000000e+00
## 183   0.0008547009 1.000000e+00 1.000000e+00
## 184   0.0000000000 1.000000e+00 1.000000e+00
## 185   0.0008591065 1.000000e+00 1.000000e+00
## 186   0.0017714792 1.000000e+00 1.000000e+00
## 187   0.0000000000 1.000000e+00 1.000000e+00
## 188   0.0000000000 1.000000e+00 1.000000e+00
## 189   0.0097087379 1.000000e+00 1.000000e+00
## 190   0.0000000000 1.000000e+00 1.000000e+00
## 191   0.0000000000 1.000000e+00 1.000000e+00
## 192   0.0000000000 1.000000e+00 1.000000e+00
## 193   0.0093457944 1.000000e+00 1.000000e+00
## 194   0.0000000000 1.000000e+00 1.000000e+00
## 195   0.0000000000 1.000000e+00 1.000000e+00
## 196   0.0000000000 1.000000e+00 1.000000e+00
## 197   0.0088495575 1.000000e+00 1.000000e+00
## 198   0.0000000000 1.000000e+00 1.000000e+00
## 199   0.0000000000 1.000000e+00 1.000000e+00
## 200   0.0000000000 1.000000e+00 1.000000e+00
## 201   0.0000000000 1.000000e+00 1.000000e+00
## 202   0.0000000000 1.000000e+00 1.000000e+00
## 203   0.0000000000 1.000000e+00 1.000000e+00
## 204   0.0000000000 1.000000e+00 1.000000e+00
## 205   0.0056022409 1.000000e+00 1.000000e+00
## 206   0.0050890585 1.000000e+00 1.000000e+00
## 207   0.0023923445 1.000000e+00 1.000000e+00
## 208   0.0000000000 1.000000e+00 1.000000e+00
## 209   0.0025252525 1.000000e+00 1.000000e+00
## 210   0.0024875622 1.000000e+00 1.000000e+00
## 211   0.0000000000 1.000000e+00 1.000000e+00
## 212   0.0000000000 1.000000e+00 1.000000e+00
## 213   0.0000000000 1.000000e+00 1.000000e+00
## 214   0.0024154589 1.000000e+00 1.000000e+00
## 215   0.0029498525 1.000000e+00 1.000000e+00
## 216   0.0000000000 1.000000e+00 1.000000e+00
## 217   0.0026455026 1.000000e+00 1.000000e+00
## 218   0.0000000000 1.000000e+00 1.000000e+00
## 219   0.0000000000 1.000000e+00 1.000000e+00
## 220   0.0000000000 1.000000e+00 1.000000e+00
## 221   0.0000000000 1.000000e+00 1.000000e+00
## 222   0.0038314176 1.000000e+00 1.000000e+00
## 223   0.0000000000 1.000000e+00 1.000000e+00
## 224   0.0000000000 1.000000e+00 1.000000e+00
## 225   0.0000000000 1.000000e+00 1.000000e+00
## 226   0.0000000000 1.000000e+00 1.000000e+00
## 227   0.0032786885 1.000000e+00 1.000000e+00
## 228   0.0000000000 1.000000e+00 1.000000e+00
## 229   0.0000000000 1.000000e+00 1.000000e+00
## 230   0.0031545741 1.000000e+00 1.000000e+00
## 231   0.0000000000 1.000000e+00 1.000000e+00
## 232   0.0040160643 1.000000e+00 1.000000e+00
## 233   0.0071428571 1.000000e+00 1.000000e+00
## 234   0.0000000000 1.000000e+00 1.000000e+00
## 235   0.0000000000 1.000000e+00 1.000000e+00
## 236   0.0000000000 1.000000e+00 1.000000e+00
## 237   0.0000000000 1.000000e+00 1.000000e+00
## 238   0.0060362173 1.000000e+00 1.000000e+00
## 239   0.0019569472 1.000000e+00 1.000000e+00
## 240   0.0000000000 1.000000e+00 1.000000e+00
## 241   0.0060240964 1.000000e+00 1.000000e+00
## 242   0.0009930487 1.000000e+00 1.000000e+00
## 243   0.0010235415 1.000000e+00 1.000000e+00
## 244   0.0020080321 1.000000e+00 1.000000e+00
## 245   0.0000000000 1.000000e+00 1.000000e+00
## 246   0.0039370079 1.000000e+00 1.000000e+00
## 247   0.0000000000 1.000000e+00 1.000000e+00
## 248   0.0010515247 1.000000e+00 1.000000e+00
## 249   0.0020366599 1.000000e+00 1.000000e+00
## 250   0.0000000000 1.000000e+00 1.000000e+00
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## 995   0.0083333333 1.000000e+00 1.000000e+00
## 996   0.0000000000 1.000000e+00 1.000000e+00
## 997   0.0000000000 1.000000e+00 1.000000e+00
## 998   0.0000000000 1.000000e+00 1.000000e+00
## 999   0.0000000000 1.000000e+00 1.000000e+00
## 1000  0.0000000000 1.000000e+00 1.000000e+00
## 1001  0.0000000000 1.000000e+00 1.000000e+00
## 1002  0.0000000000 1.000000e+00 1.000000e+00
## 1003  0.0000000000 1.000000e+00 1.000000e+00
## 1004  0.0000000000 1.000000e+00 1.000000e+00
## 1005  0.0000000000 1.000000e+00 1.000000e+00
## 1006  0.0000000000 1.000000e+00 1.000000e+00
## 1007  0.0000000000 1.000000e+00 1.000000e+00
## 1008  0.0000000000 1.000000e+00 1.000000e+00
## 1009  0.0000000000 1.000000e+00 1.000000e+00
## 1010  0.0000000000 1.000000e+00 1.000000e+00
## 1011  0.0000000000 1.000000e+00 1.000000e+00
## 1012  0.0000000000 1.000000e+00 1.000000e+00
## 1013  0.0000000000 1.000000e+00 1.000000e+00
## 1014  0.0000000000 1.000000e+00 1.000000e+00
## 1015  0.0088495575 1.000000e+00 1.000000e+00
## 1016  0.0000000000 1.000000e+00 1.000000e+00
## 1017  0.0000000000 1.000000e+00 1.000000e+00
## 1018  0.0000000000 1.000000e+00 1.000000e+00
## 1019  0.0000000000 1.000000e+00 1.000000e+00
## 1020  0.0000000000 1.000000e+00 1.000000e+00
## 1021  0.0000000000 1.000000e+00 1.000000e+00
## 1022  0.0000000000 1.000000e+00 1.000000e+00
## 1023  0.0000000000 1.000000e+00 1.000000e+00
## 1024  0.0000000000 1.000000e+00 1.000000e+00
## 1025  0.0000000000 1.000000e+00 1.000000e+00
## 1026  0.0000000000 1.000000e+00 1.000000e+00
## 1027  0.0000000000 1.000000e+00 1.000000e+00
## 1028  0.0000000000 1.000000e+00 1.000000e+00
## 1029  0.0000000000 1.000000e+00 1.000000e+00
## 1030  0.0000000000 1.000000e+00 1.000000e+00
## 1031  0.0000000000 1.000000e+00 1.000000e+00
## 1032  0.0084033613 1.000000e+00 1.000000e+00
## 1033  0.0000000000 1.000000e+00 1.000000e+00
## 1034  0.0000000000 1.000000e+00 1.000000e+00
## 1035  0.0000000000 1.000000e+00 1.000000e+00
## 1036  0.0076335878 1.000000e+00 1.000000e+00
## 1037  0.0000000000 1.000000e+00 1.000000e+00
## 1038  0.0000000000 1.000000e+00 1.000000e+00
## 1039  0.0000000000 1.000000e+00 1.000000e+00
## 1040  0.0000000000 1.000000e+00 1.000000e+00
## 1041  0.0000000000 1.000000e+00 1.000000e+00
## 1042  0.0000000000 1.000000e+00 1.000000e+00
## 1043  0.0000000000 1.000000e+00 1.000000e+00
## 1044  0.0000000000 1.000000e+00 1.000000e+00
## 1045  0.0000000000 1.000000e+00 1.000000e+00
## 1046  0.0000000000 1.000000e+00 1.000000e+00
## 1047  0.0000000000 1.000000e+00 1.000000e+00
## 1048  0.0076923077 1.000000e+00 1.000000e+00
## 1049  0.0073529412 1.000000e+00 1.000000e+00
## 1050  0.0000000000 1.000000e+00 1.000000e+00
## 1051  0.0000000000 1.000000e+00 1.000000e+00
## 1052  0.0000000000 1.000000e+00 1.000000e+00
## 1053  0.0000000000 1.000000e+00 1.000000e+00
## 1054  0.0000000000 1.000000e+00 1.000000e+00
## 1055  0.0125000000 1.000000e+00 1.000000e+00
## 1056  0.0000000000 1.000000e+00 1.000000e+00
## 1057  0.0000000000 1.000000e+00 1.000000e+00
## 1058  0.0000000000 1.000000e+00 1.000000e+00
## 1059  0.0000000000 1.000000e+00 1.000000e+00
## 1060  0.0000000000 1.000000e+00 1.000000e+00
## 1061  0.0000000000 1.000000e+00 1.000000e+00
## 1062  0.0000000000 1.000000e+00 1.000000e+00
## 1063  0.0000000000 1.000000e+00 1.000000e+00
## 1064  0.0000000000 1.000000e+00 1.000000e+00
## 1065  0.0000000000 1.000000e+00 1.000000e+00
## 1066  0.0000000000 1.000000e+00 1.000000e+00
## 1067  0.0000000000 1.000000e+00 1.000000e+00
## 1068  0.0000000000 1.000000e+00 1.000000e+00
## 1069  0.0000000000 1.000000e+00 1.000000e+00
## 1070  0.0000000000 1.000000e+00 1.000000e+00
## 1071  0.0000000000 1.000000e+00 1.000000e+00
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## 1073  0.0112359551 1.000000e+00 1.000000e+00
## 1074  0.0000000000 1.000000e+00 1.000000e+00
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## 1076  0.0000000000 1.000000e+00 1.000000e+00
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## 1078  0.0000000000 1.000000e+00 1.000000e+00
## 1079  0.0000000000 1.000000e+00 1.000000e+00
## 1080  0.0000000000 1.000000e+00 1.000000e+00
## 1081  0.0108695652 1.000000e+00 1.000000e+00
## 1082  0.0000000000 1.000000e+00 1.000000e+00
## 1083  0.0000000000 1.000000e+00 1.000000e+00
## 1084  0.0000000000 1.000000e+00 1.000000e+00
## 1085  0.0102040816 1.000000e+00 1.000000e+00
## 1086  0.0000000000 1.000000e+00 1.000000e+00
## 1087  0.0000000000 1.000000e+00 1.000000e+00
## 1088  0.0000000000 1.000000e+00 1.000000e+00
## 1089  0.0000000000 1.000000e+00 1.000000e+00
## 1090  0.0000000000 1.000000e+00 1.000000e+00
## 1091  0.0000000000 1.000000e+00 1.000000e+00
## 1092  0.0000000000 1.000000e+00 1.000000e+00
## 1093  0.0109890110 1.000000e+00 1.000000e+00
## 1094  0.0000000000 1.000000e+00 1.000000e+00
## 1095  0.0000000000 1.000000e+00 1.000000e+00
## 1096  0.0000000000 1.000000e+00 1.000000e+00
## 1097  0.0000000000 1.000000e+00 1.000000e+00
## 1098  0.0000000000 1.000000e+00 1.000000e+00
## 1099  0.0000000000 1.000000e+00 1.000000e+00
## 1100  0.0000000000 1.000000e+00 1.000000e+00
## 1101  0.0000000000 1.000000e+00 1.000000e+00
## 1102  0.0000000000 1.000000e+00 1.000000e+00
## 1103  0.0000000000 1.000000e+00 1.000000e+00
## 1104  0.0000000000 1.000000e+00 1.000000e+00
## 1105  0.0000000000 1.000000e+00 1.000000e+00
## 1106  0.0000000000 1.000000e+00 1.000000e+00
## 1107  0.0000000000 1.000000e+00 1.000000e+00
## 1108  0.0000000000 1.000000e+00 1.000000e+00
## 1109  0.0000000000 1.000000e+00 1.000000e+00
## 1110  0.0000000000 1.000000e+00 1.000000e+00
## 1111  0.0000000000 1.000000e+00 1.000000e+00
## 1112  0.0076335878 1.000000e+00 1.000000e+00
## 1113  0.0000000000 1.000000e+00 1.000000e+00
## 1114  0.0000000000 1.000000e+00 1.000000e+00
## 1115  0.0000000000 1.000000e+00 1.000000e+00
## 1116  0.0000000000 1.000000e+00 1.000000e+00
## 1117  0.0000000000 1.000000e+00 1.000000e+00
## 1118  0.0000000000 1.000000e+00 1.000000e+00
## 1119  0.0000000000 1.000000e+00 1.000000e+00
## 1120  0.0000000000 1.000000e+00 1.000000e+00
## 1121  0.0000000000 1.000000e+00 1.000000e+00
## 1122  0.0000000000 1.000000e+00 1.000000e+00

top.bulk.mods <- head(MOD.dt, 10)$Module1.name %>% unique()
# One hub per module
bulk.hubs <- data.table::fread("./Data/Katia_Modules/hubs_gencode30.tbl.txt")
top.bulk.hubs <- subset(bulk.hubs, module %in% top.bulk.mods) 
# Plot hub expression in sc data
monocle3::plot_cells(cds,
                     genes=top.bulk.hubs$symbol,
                     group_cells_by="cluster",
                     color_cells_by="cluster",
                     show_trajectory_graph=F) #  rasterize = T (for super high-res figures) 

Check c